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S DEPARTMENT OF THE INTERIOR S GEOLOGICAL SURVEY ATIONAL WETLANDS RESEARCH CENTER 30 CAJUNDOME BLVD 9FAYETTE LA 70506-31 52 BY Samuel M. Carney Washingon, D.C. 1992 U.S. Department of the Interior U.S. Fish and Wildlife Service P rocedures have been developed over the past 30 earsf for managing the take of waterfowl by hunters. Wings of ducks contributed voluntarily to the U. S. Fish and Wildlife Service by hunters are examined each year by experts. This procedure involves the determination of species, sex, and age of ducks through an examination of these detached wings. Many persons skilled at examining wings of ducks have retired in recent years. We became concerned that these skills might be lost to Jitture generations ij'not properly documented. Documenting such skills is dificult. Subtle dzferences in feather color andfeather texture are used to distinguish young ducks from adult ducks and males from females. Printed words and pictures have their limitations, but we hope this publication captures the technique and preserves it. /' Director U.S. Fish and Wildlife Service Introduction .......................................................... 1 Feather groups of the duck wing ............................ 2 Mallard .................................................................. 5 American black duck ........................................ 1 O Mottled duck ....................................................... 13 Gadwall .............................................................. 1 6 American wigeon ................................................. 19 Green-winged teal ............................................... 24 Blue-winged and cinnamon teals ......................... 28 Northern shoveler ................................................ 3 1 Northern pintail .................................................. 34 Wood duck .......................................................... 39 Harlequin duck ................................................... 42 Steller's eider ........................................................ 45 Separation of redhead and canvasback ................. 48 Redhead .............................................................. 49 Canvasback. ......................................................... 54 Separation of greater and lesser scaups ................. 59 Greater scaup ....................................................... 59 Lesser scaup ......................................................... 6 5 Ring-necked duck ............................................... 70 Separation of common and Barrow's goldeneyes ......................................... 75 Common goldeneye .........................................7..5. Barrow's goldeneye ...........................................-.8. 1 Separation of bufflehead and hooded merganser .........................................8..6 Bufflehead ........................................................8..7. Hooded merganser ............................................9..2 Separation of red-breasted and common mergansers .....................................9..6 Red-breasted merganser ......................................9. 6 Common merganser .......................................... 100 Separation of oldsquaw. black scoter. and surf scoter .............................................. 0 5 Oldsquaw .......................................................1..0.6 Black scoter ....................................................... 1 0 Surf scoter ......................................................... 114 White-winged scoter ........................................ 118 Common eider ............................................. 122 King eider .......................................................... 127 Ruddy duck ....................................................... 132 Fulvous whistling duck ...................................... 135 Black-bellied whistling duck .............................. 138 Appendix: Key to duck species .......................... 141 T his publication contains procedures used by the U.S. Fish and Wildlife Service to determine the species, sex and age composition of the harvest of North American ducks using detached wings contributed by hunters. Original studies of the use of duck wings for this purpose began in 1958 and were led by the author and A.D. Geis. Others con-tributing to these studies include R.L. Croft, E.M. Martin, A.N. Novara, L.D. Schroeder, M.G. Smart, and M.F. Sorensen. Major suppliers of known-age specimens include: W. Anderson, R.S. Billard, A.J.Erskine, D. Hall, A.S. Hawkins, C. Hoffpauir, L.R. Jahn, R. L. Jessen, F.B. Lee, J.J. Lynch, R.K. Martinson, D.I? Olson, R.I? Osbolt, C. Ritcey, R.N. Smith, H.E. Spencer, V.D. Stotts, and J. Takekawa. R.I. Smith edited the text, A.J. Godin prepared Figure 2, Larry Ketchum Photography took the pictures, and R.E. Cummins typed the manuscript. The Waterfowl Parts Survey became the means by which large samples of duck wings were obtained. This survey became national in scope in 1961. Collection of goose tails was added to the survey in 1962. Packages of envelopes are mailed to selected hunters who return wings from shot ducks and tail feathers from shot geese by mail to collection points throughout the United States where they are exam-ined to determine species, sex, and age. For readers who are interested in more detailed information on the development and testing of procedures described in the following pages, copies of a more technical report, which was prepared by the author, are available by writing to Waterfowl Harvest Surveys, U.S. Fish and Wildlife Service, 10800 Laurel-Bowie Road, Laurel, Maryland 20708-3600. That report, titled Observations on Sexing and Aging Ducks Using Wings, makes several points that must be understood by anyone who plans serious use of these techniques. The proce-dures presented here vary among species in the degree to which they accurately identify the age and sex of duck wings. Accuracy varies according to skills of the observer. Some individuals become highly skilled after examining large numbers of duck wings, while other individuals never develop the ability to detect subtle differences in feather texture and feather quality with high levels of profi-ciency. Levels of skill will decline if techniques are not practiced with regularity. Terminology related to age must be clarified. An immature wing possesses one or more characteristics known to be associated with ducks hatched in the most recent nesting season. Since the term imma-ture often refers to maturing processes not related to plumage, juvenalmight have been more appropri-ate. Nevertheless, after 30 years of use in this con-text, introducing another age designation would serve no purpose. All wings not possessing charac-teristics associated with the most recent hatch are classified as adult except those of male eiders which are identified as being from sub-adults. To determine the species, age, and sex of ducks from detached wings, a worker must be familiar with the various feather groups (Figure 1). The first step is to determine the species represented. A key to species is included as an appendix to this publica-tion. Usually, slight differences in feather shape, color, pattern, wear, or replacement are sufficient during the fall and winter to separate immatures from adults. Age determination is a step-by-step search for one or more traces of immature plumage. Wings on which no traces of immaturity can be found, or in some cases those that have positive adult characters, are considered to be from adults. During their first fall and winter immatures of many of the more common species of ducks molt certain wing-feather groups located near the body and replace them with adult-type feathers. Those feathers that are replaced include the tertials, greater tertial coverts, post humerals, and scapulars. Scapulars are of limited use in classifying wings because most hunters do not include scapulars on the wings they remove. Tertials, as defined here, are actually the more proximal secondaries, which are generally different in size, shape, and color from their more distal counterparts. These feathers are often sexually dimorphic and usually molt with adjacent body feathers. Post humerals are feathers attached to the humerus. They lie between the tertials and scapulars. They usually molt with adja-cent body feathers. Greater coverts are the first row continued on page 4 FEATHER GROUPS OF THE DUCK WING REMIGES: (Flight feathers) Alula: The feathered "thumb" of the bird wing Primaries: Flight feathers attached to the hand (manus) Secondaries: Flight feathers attached to the forearm (ulna) Tertials: Incorrect (morphologically) but used here to designate the more prox-imal secondaries which are generally different in size, shape, and color from their more distal counterparts, are often sexually dimorphic, and usually molt with adjacent body feathers. Post humerals: Feathers attached to the humerus. They lie between the tertials and scapulars, usually molting with adjacent body feathers. Scapulars: Feathers of the humeral (upper arm) feather tract. These lie on either side of the back and may partially cover a folded wing. They usually molt with adjacent body feathers. (Not shown) Axillars: Elongate feathers growing in the "armpit" region and closing the space between the spread wing and the body WING COVERTS: (Cover flight feathers) Greater coverts: The first row of feathers overlying the flight feathers, identified by the particular feathers they cover as primary, secondary, or tertial coverts. Tertial coverts: Those greater coverts that overlie the tertials. Designated separately here because they are sexually dimorphic in adults of several species and usually molt with the adjacent body feathers. Middle coverts: The next row of coverts. Lesser coverts: The next several rows of coverts. Marginal coverts: An indefinite number of rows anterior to the lesser coverts 2 PRIMARIES Figure 1. Feather groups of a typical dabbling duck wing of feathers overlying the flight feathers, identified by both fresh wings and wings with varying degrees of the particular feathers they cover as primary, sec- stiffness, as they are commonly received through the ondary, or tertial coverts. Greater tertial coverts are Waterfowl Parts Survey. To ensure uniformity all those greater coverts that overlie the tertials. They measurements are made using a standardized proce-are designated separately because they are sexually dure and measuring board (Figure 2). Measure-dimorphic in adults of several species and often ments are referred to as wing notch-length. molt with the adjacent body feathers. The degree to Tables have an advantage over keys in that the which these feathers are replaced is quite variable, color, shape, or texture of a particular group of I even among closely related species. Southern nest- feathers can be compared on one page across the ing duck species may initiate upperwing molts in four age and sex categories. Unlike keys, tables do the fall rather than in the spring. Fall wing molting not lead one directly to the answer. Despite this occurs among wood ducks, mottled ducks, and limitation, most people prefer tables to keys. whistling ducks. Such molting reduces the accuracy Therefore, tables are used in this publication to of immature wing identification in the fall and present information on individual species. Wing characteristics are not always listed in the same Male wings on most North American ducks are sequence for each species. They are listed in the slightly larger than those of females. For a few sequence in which they can be most efficiently used. species, this difference is large enough to permit A brief narrative, which identifies the most separation of the sexes using wing measurements. frequently used wing characters in a table, accompa- The procedure for measuring duck wings applies to nies most tables. Figure 2. Procedure for measuring wings 4 Place the heel of one hand over the end of measuring board and grasp wing in the area of the radius and ulna. Pull until wooden block seats tightly into the notch at the bend of the wing. Flatten the leading edge of the wing and note the length of the wing at the longest primary. Note: If primary quills are not completely grown, the measurernent is not val~d. Sex Determination The white bar anterior to the speculum extends onto the greater tertial coverts on all female wings but terminates at the proximal edge of the specu-lum on nearly all male wings. Approximately 2%-3 percent of males show some white edging on their tertial coverts. Adult males can be identified because the white is not continuous with that over the sec-ondaries. Immature males with white over the ter-tial coverts are difficult to tell from immature females. The white bar is the easiest sex character to use, because of its high degree of reliability and the fact that it is rarely lost when a wing is detached. Vermiculated scapulars are found only on males. Early in the hunting season (September and October) many males possess barred scapulars which are remnants of their summer plumage. Proximal underwing coverts are vermiculated or flecked on adult and most immature males. These feathers are barred on females and on a few imma-ture males. Age Determination of Males Immature tertials are often frayed and faded, usually narrow, and lack the pearly color of adult tertials. By late November immature tertials are replaced by first winter tertials, identical in appear-ance to adult feathers. At the same time, immature tertial coverts may be replaced by coverts which are broader, unfrayed, and similar to adult coverts and thus differ from adjacent immature coverts, which have not been molted. Many immature males have light edging on the inner webs of the most distal primary coverts. Adult males do not show this char-acter. Middle coverts of immatures are narrower and more trapezoidal than those of adults. Occasionally, these feathers on immature males are worn and/or have light edges. This type of edging does not occur on the middle coverts of adult males. Age Determination of Females Tertials that are frayed and/or faded are remnants of immature plumage and are found only on imma-ture ducks. Tertial coverts of immatures are often frayed, faded, and narrow, and the two most proxi-mal often lack the white of the speculum bar. As with males, greater tertial coverts of immatures may be replaced. Conspicuous light edging on the inner webs of the four most distal primary coverts is found only on immatures. Adults may have minute or no edging on these coverts. ~ i d d l ceo verts of immatures tend to be narrow and trapezoidal, while those of adults are broadly rounded. Scapulars 1 Venniculated o'barred or both / Barred + -. - I Wing Character I Proximal Flecked to vermiculated I Barred 1 I Male 1 Female Adult 1 Immature / Immature 1 Adult Tertials - - - - - - roiinded;.sotid gray to brown, no edgtljs Broad, pearly gray, no edging, rarely frayed or faded' May be growing in October Primary coverts Small, narrow, brownish, often frayed or faded near tips. Adul type feathers may be growing in December or later After mol: Similar to adul After mol: Similar to adul male female Broadty rounded; brown; ~suaHyw Sth light brown @iQlng; sometimes no SiigMfy ttiangukr to W o i d a l ; often frayed or faded; usually bmwn No edging Varies from pearly gray to brownish, often light edging; rarely frayed or faded Ma) have fiie'edging or no Edging varies from w9 Inner web of four most distal have llght edg~ngo r no edging conspicuous to absent edging I Inner web of four most distal wlthout edglng to conspicuous edging Innerweb of four most distal wlth llght edglng or no edging I m.; a IFigur-e 5. Adult hniale rnallrird ~ v i r hu necigid co\.ct-ts Figure 6. Immature male mallard with immature tertials Age Determination Sex Determination of Adults Because wings of the sexes are similar, it is easier Adult male tertials are more than 90 mm. long to determine the age of black ducks before attempt- from the edge of the longest tertial covert to the ing to identify their sex. Tertials small, narrow, and tertial tip, and acutely pointed with some pearly frayed andlor faded near their tips are remnants of color on the outer webs. Tertial coverts have broad immature plumage. Adult tertials are longer and edging which is pale brown. Middle and lesser wider and are not frayed or faded. Tertial coverts of coverts are broadly rounded and unfrayed. The the immature plumage tend to be narrow, some- notch-length of 94 percent of the adult male what trapezoidal, and frequently frayed andlor known-age specimens was greater than 28 1 mm. faded. Those of adults are wide, broadly rounded, Adult female tertials are less than 90 mm. from and rarely frayed or faded. During their first fall and the edge of the longest tertial covert to the tertial winter, a substantial fraction of the immatures may tip, and they are rather bluntly pointed. Pearly color replace both their immature tertials and tertial generally does not occur on the outer webs. Tertial coverts with adult-type feathers. Thus, it is impor- coverts have broad edging which is pale brown. tant to look closely at the middle coverts immedi- Middle and lesser coverts are broadly rounded and ately anterior to the tertial coverts for indications of unfrayed. The notch length of 94% of adult females the trapezoidal shape, duller color, and wear that was less than 281 mm. indicate immaturity. Primary coverts of many Immature tertials less than 88 mm. from the immatures have light edging on their inner webs. longest covert to the tertial tip are from females and This type of edging does not occur on adults. longer tertials are from males. 1 Wino 1 Male 1 Female 1 I Y"Y'YY'"' I Adult I Immature I Immature I Adult I 1 After molt: Similar to adult After molt: Similar to adult rounded'arc; not frayed of male female 1 faded 1 Greater 1 Four most distal do not have 1 Four most distal otten (but not always) have light edging on 1 Four most distal do not have I ure 11. Adult female black duck p n p y3qq spur a~naewwI' ~ ;a1rnS !j - - 1 Because mottled ducks are southern nesters, most females, but there is considerable overlap. Birds immatures have replaced both tertials and greater with three or more non-iridescent secondaries are tertial coverts at the time hunting occurs. Careful virtually all females, but birds with only one or two scrutiny, however, will usually reveal one or more non-iridescent secondaries may be of either sex. faded coverts in this area. These are remnants of the The greater tertial coverts of females tend to more immature plumage. Sexual identification is difficult. heavy edging than those of males, but the two types Wings of males are generally longer than those of grade together. Male I Female , Adult 1 Immature I Immature I Adult Similar to adult males; lack Similar to aduk females; I Often bluntly pointed; internal markings oflen internal markings often r mm.; lack internal . .- 4 Broadly rounded; edging Usually similar to those of Usually similar to those of Broadly rounded with heavy varies from broad to narrow adult males; often one or adult females; sometimes tan edging more narrow and frayed or one or more narrow and faded fraved or faded 1 Notch-length I 81% > 255 mm. 1 84% > 251 mm. 1 81% < 250 mm. 1 86% < 254 mm. 1 Sex Determination of Adults Greater, middle, and some lesser coverts of adult males are mostly either black or cinnamon. On adult females, black and cinnamon feathers are much restricted to the posterior three or four rows of coverts. Tertials of adult males are long, acutely pointed, silver-gay without edging or tipping. Those of adult females are much shorter, more bluntly pointed, and silver-brown with cream col-ored tips. Greater tertial coverts of adult males are part black and part gray, rarely with traces of white tipping. Those of adult females are similar but usu-ally well tipped with white. Marginal coverts of adult males are without edging but have arcs of narrow vermiculation. On adult females, these feathers are similar to the lesser coverts and ofcen have edging but may be either plain or with wide internal bars or arcs. Post humerals of adult females usually have cream edging at their tips. Those of other ages and sexes do not have this edging. Sex Determination of Immatures Tertials of both sexes are short, bluntly pointed, . . and ofieti frayed at their tips. They are quite similar to those of adult females. In late fall, they may be replaced by sexually dimorphic adult-type tertials. Greater tertial coverts of both sexes usually appear part black and part gray and are tipped with cream. They are narrower and more pointed than those of adults. Greater, middle, and sbme lesser coverts of immature males have some black andlor cinnamon in three or more rows. Immature females usually have little or no cinnamon color and black is ofien restricted to two rows of coverts. Some males show arcs of narrow vermiculation. Both sexes may show pale barring, which is usually wider on females. Notch-length measurements are usehl to identify immatures by sex, as long as the shafts of the pri-mary feathers have hardened. In 90 percent of the specimens measured, immature male notch-lengths were equal to or greater than 255 mm., while those of immature females were less than 255 mm. --- ------- Immature Immature 1 -- - Scapulars v o r b a n e d o r b a t h - - . -- - -- -- . I Barred Tertials Long, acutely pointed; Small; brownish; bluntly pointed; usually Shorter; bluntly pointed; silver-gray; without edging; ------ gray-brown with tips not hayed Or faded cream-colored tips which are not frayed or faded Greater tertial Bluntly pointed; outer webs Somewhat pointed; outer webs black; inner webs brown; Broadly rounded; usually coverts black; inner webs gray, often frayed and faded at tips with white tipping; outer 1 rarely with traces of white webs black or brown; inner tipping; not frayed or faded k - 7 After molt: Similar to adult After molt: Similar to adult fwaedbesd brown; not frayed or 1 Post 1 Broad, rounded, tips without 1 Narrow; pointed; tips without edaino 1 Broad rounded tips, usuallv 1 1 humerals edaing 1 1 lia- ht eda-ina" . I h- o.t z n a t h 7 5 e 2 6 2m m.. .. . . I 80% > 255 -mm. .. ( 80#s 255mm. I 9 ~ % c ~ r n m . 1 Adult males have a large white uppenving patch, long acutely pointed tertials with black outenvebs, and gray greater tertial coverts that are somewhat pointed and narrowly white-edged. Adult females have a sharply defined white edging on both greater and middle tertial coverts. Usually the sharp white edging on the middle and lesser coverts is sufficient to identify adult females, however, a few are so lightly colored as to resemble immature males. A carehl check of tertials and greater tertial coverts will serve to identify them. Immature males usually have small, brownish tertials and tertial coverts, but by November these are often replaced with adult male-type feathers. The middle and lesser coverts are gray-brown and indistinctly edged with a lighter gray. Immature females have small, brownish tertials and tertial coverts similar to those of immature males. By the middle of the fall hunting season these may be replaced with adult female-type feath-ers. The middle and lesser coverts are brownish with well-defined pale tan edges. Often the greater secondary coverts lack most of the black tipping common to wings of other ages and sexes and their outer webs are brownish gray to gray-white. Separation of American wigeon wings from Eurasian wigeon wings can be accomplished using the following procedures: American wigeon Axillars (if present) entirely white or flecked only at their tips. Undenving middle coverts all or nearly all white. Scapulars of males (if present) are heavily vermic-ulated with reddish brown. Eurasian wigeon Axillars (if present) heavily flecked with gray over their entire length. Underwing middle coverts heavily flecked with gray. Scapulars of males (if present) are heavily vermiculated with black and white. 1 Male KYaeter 1 Adult 1 Immature 1 Female I Immature I ~ d u ~ t I . -- Scapular~ I Vmiculated w barred or both -- I Barred A. Underwing Coverts gray, heavily flecked with white Coverts brownish; heavily edged with white to gray ( Occasionally as in females I Brown with light gray-brown edging; often frayed and faded Dark brown; edged with After molt: Similar to adult white to form a broadly I narrow white edging After molt: Similar to adult rounded arc male female GREEN-WINGEDT EAL If the scapulars are attached, the presence of one frayed. Adult female-type tertials have cream col-or more vermiculated feathers indicates that the ored edging. Adult male-type tertials are long, with-wing is from a male. If none of the scapulars are out edging, or with narrow edging. The presence of vermiculated, the wing could be that of either a adult-type tertials is not by itself an indication of male or female. age. Many geen-winged teal molt tertials during The stripe on the most distal tertial is the most the hunting season. For some of these, it is possible useful indicator of sex. On the wings of males, this to split incoming pin feathers to see whether the stripe is black and sharply delineated from the basic new feather is male or female in character. For feather color. On the wings of females, this stripe is wings with molted tertials, determination of sex is black to brown, but grades into the basic feather not possible. color. The sex of approximately half of the imma- Tertial coverts that are long and narrow, showing ture birds can be identified by measurements. fine, light edging, and a frayed fringe are remnants Wings 183 mm. or longer are from males and of immature plumage and positively identify the wings shorter than 175 mm. are from females, but wing as being immature. The tertial stripe may be sex cannot be determined from measurements used to separate the sexes. Adult female-type tertial between 176 mm. and 182 mm. A sample of wings coverts are broadly rounded with wide edging that from Adak Island, Alaska, green-winged teal (con- is cream or brown. Adult male-type tertial coverts sidered a different subspecies) averaged approxi- are a uniform gray, sometimes with a narrow bu@ mately 5 percent longer than wings collected in the edging, and tapering to a blunt point. Adult-type lower 48 states. The number of iridescent secon- tertial coverts are not a reliable indicator of age. daries differs between sexes. Wings having fewer Middle coverts of adult males are smooth and than four secondaries with completely iridescent without edging. Middle coverts of immatures are green outer webs are usually from females and rough and may show gray-appearing wear at their wings with five or more such secondaries are usually edges. Immatures may also possess light edging from males. Sex cannot be determined in this man- which is more pronounced on females, but may also ner from wings with four or four and a half irides- be present on males. Broad, rounded middle coverts cent secondaries. with wide edging are found only on adult females. Immature tertials are small, narrow, and rather Traces of immature plumage usually remain most delicate. The tips of these feathers are often badly obvious immediately anterior to the tertial coverts. Wing Male 1 Female Character Adult I Immature 1 Immature I Adult Tertials Uniform gray; tapering to a Small; narrow; rather delicate tips are often badly frayed; Buff edging including usually buff edging rounded tips; unfrayed; longitudinal stripe often poorly defined along inner edge; may be brown or black; stripe sometimes well defined; may be molting 1 1 ( male 1 female I I 1 Middle and Broadly rounded; no edging; Gray with wear around edges; appear ragged; somewhat Broadly rounded; usually lesser coverts gray that matches tertial narrow and trapezoidal; late in year contrast to a variable with wide light edgjng but coverts deoree with new freolaced) oreater coverts sometimes no eda~na Sex Determination The speculum of males is a bright iridescent green and that of females is a dull non-iridescent green. Rarely, a female may show a trace of irides-cence. The greater secondary coverts of males appear entirely white on their outer webs while those of females are heavily dark spotted and fre-quently appear more dark than light. A few males may show some spots in the greater coverts. Age Determination of Males Tertials of adults are greenish black, rarely frayed, andyery long and pointed. They are trimmed with a very narrow tan edging. Tertials of the immature plumage have wide edging which is tan and they are much shorter and more bluntly pointed than those of adults. They are brownish and often somewhat frayed and faded at the tips. Adult-type tertials occur on immatures and are similar to those of adult males but are usually still growing in December. Adult and replaced tertial coverts of immatures are similar, i.e. a dark brownish black without edging but usually showing a blue wash. Immature-type tertial coverts are brown, usually with pronounced edging and tan and usually not washed with blue. Age Determination of Females Tertials of adults are much more bluntly pointed than are the immature tertials. In addition, imma-ture tertials frequently become somewhat frayed at their tips. Tertial coverts of adults are usually straight sided to a rounded tip, whereas those of immature birds usually narrow slightly to a rounded tip that often shows traces of fading. Well propor-tioned white inverted "V's" generally indicate adults. -- -- --- , Immature I Adult Greater coverts Tertial Broadly rounded; dark coverts brown washed with blue; brown usually with usually without edging; not pronounced light brown frayed or faded I female The presence of white primary shafts separates shovelers from all other North American ducks. All birds having specula that are more than half gray or dull non-iridescent green are females. All males and a few adult females have iridescent green on more than half of their secondaries. Most females show cream edging on the lesser and middle coverts. This often covers all these feathers, particularly on adult females. Immature males may show a few cream colored edges on feathers near the alula. The tertials of immature males are brownish and much frayed, while those of adults and first winter immatures are greenish black and much longer. The tertial covcrts of the immature male are brownish black and often show a frayed fringe. Adult tertial coverts are blackish, often washed with blue. Both immature tertids and immature tertial coverts are usually present during October. Immature males generally have small dusky spots on their greater coverts, while adults do not. The immature tertials of females are similar to those of immature males. The tertials of adult females are wider, not frayed, and more heavily washed with white at the tips. Male I Female EFacIer 1 Adul, I Immature 1 Immature I Adult Smoothly rounded; dark Brown; often much frayed; with trace of light edging Brown; white edging forms brown to shiny black; After molt: Similar to adult After molt: Similar to adult smoothly rounded arc unfrayed male female White; dark bases normally White; often with small dark White; about 114 - 112 inch of dark base normally shows covered by middle coverts spots at tip; about 114 - 112 lnch of dark base normally shows 31 ' - Sex Determination one wing in 20 shows faint light edging, but the The speculum is at least partly iridesceRgreen general aspect of these is such that they can readily (rarely purple) on males and a dull non-iridescent be identified as adults. F bronze or rarely non-iridescent green on females. The tertial coverts of the immature plumage have The underwings of most males are flecked to ver- conspicuous edging which is a light yellowish miculated; those of a few males and all females are brown. Birds with such wings are immature males. barred. Male adult and first winter tertials are long Tertial coverts of the adult-type lack edging and are and gray and the innermost has a wide marginal found on both adult and immature wings. Tertials black stripe. The tertials of females are shorter and that lack the black edging are immature. It seems more brownish and lack the black stripe. Immature lilcely that wings with new tertials growing after tertials of both sexes are similar and somewhat like November are immatures. those of adult females. Vermiculated scapulars are found only on males, but unvermiculated sca~ulars Age Determination of Females may be present on both males and females. The middle coverts of the immature plumage are Tertials and scapulars are useful for identibing rather narrow and somewhat trapezoidal. Any bar-some males, but not for separating the sexes in all ring of these feathers is largely made up of triangu-cases. lar patches at the feather edge. The middle coverts of adults are broad and rounded and any barring is Age Determination of Males made up of variable shaped patches recessed from The most useful character for separating the ages the feather's edge. is the condition of the middle coverts. These have The tertial coverts of the immature plumage are light edging on immatures and often appear narrow, very frayed, often poorly edged and individual and frayed. Some of these feathers persist through- feathers are often long and narrow. The tertial out the hunting season. On adult males, these coverts of adult-type plumage are not very frayed, feathers are not frayed and generally plain. About are well rounded, and have conspicuous edging. Wing Male I Female Character 1 I Immature I Immature Adult I Underwing Gray; heavily flecked with white I Alternate brown and white bars I Tertials I Acutelv oointed: brownish I Brown: most distal with 1 Brown with oale lonaitudinal I Bluntlv oointed: dark brown: I I surrounding rachis I often frayed 1 frayed 1 brownish longitudinal stripe; I Middle and I Grav: without edaina: I Grav: liaht brown edaina at Brown with liaht edaina: 1 Brown with liaht edaina: edging; somewhat 1 from edging; rounded trapezoidal Immature f Adult Figure 42. Common middle covert markings of female northern pintail Sex Determination and extends farther distally than on wings of imma- The white trailing edge of the secondaries on tures. females is much wider on the outer webs than on the inner webs. On males, this white edge is Age Determination of Females approximately the same width on both webs. On Immature tertials are pale bronze with pointed, females, the first secondary proximal to these white- usually frayed, tips. Immature tertial coverts are edged feathers is washed with black on the outer narrow and yellow-geen. Most immatures replace web. Males have no black on this feather. both tertials and tertial coverts with purplish red adult-type feathers by late October. Usually, the Age Determination of Males blue iridescence is confined to two rows of coverts Immature tertials are pale bronze with pointed, and is restricted to the proximal half of the imma-usually frayed, tips. Immature tertial coverts are ture wing. Adult coverts are somewhat wider and narrow and yellow-green. Most immatures replace tend to lie more smoothly on the wing. Dark blue both tertials and tertial coverts with dark blue usually extends onto the third row of coverts and adult-type feathers by late October. As the imma- extends farther distally than on the immature ture middle and lesser coverts are replaced by the coverts. adult-type plumage, the presence of a few dark blue feathers among the duller coverts indicates immatu- Note: rity. Usually, the dark blue does not extend onto the By mid-fall, some immature wood ducks hatched third row of coverts and is much restricted to the in the southern United States appear to have area anterior to the ~roximahl alf of the secondaries. replaced all of their immature upper-wing coverts. Generally, coverts of adults are somewhat wider and Their wings cannot be distinguished from those of tend to lie more smoothly on the wing and the dark adults. blue usually extends on to the third row of coverts 1 Male I Female I , , blue-black; second most distal white-edged at itst ip A--fIt-e r molt: Similar to adult LAn-..lf.. ter molt: Similar to adult rounded tips 1 Other 1 Greater and middle coverts Greater coverts and middle Greater coverts often 1 Greater coverts usually 1 anteriorly over three or more confined to two rows of to two rows of cdverts three or more rows of rows of coverts coverts nearest secondaries anterior to secondaries coverts I pa A~J) IOU pue papunod h~qloo~us sd!~: 1 ~ 061 6u1peus' y3els SEPARATIONO F REDHEAD AND CANVASBACK Adult male canvasbacks have much more white secondary coverts and secondaries is greater on on the upper surface of their wings than any other wings of redheads of all age and sex groups than it is age-sex category of redhead or canvasback. All other on the wings of adult female or immature canvas-canvasbacks have much darker secondaries than backs of either sex. those of all redheads. Thus, the contrast between Immature greater coverts are narrower, squared, often frayed to a point over the tertials with an indistinct pale tip over the secondaries. During the hunting season, immature birds occasionally replace both their immature tertials and greater tertial coverts with adult-type feathers. For a given sex, these new feathers are indistinguishable from those of adults but differ markedly from other immature greater and middle coverts which are retained. With practice, redheads can be aged primarily by the appearance of their tertial coverts. Several combina-tions are possible: 1) immature-type tertial coverts always indicate an immature bird but sex is best determined from other coverts; 2) adult male or adult female-type coverts similar to the surrounding coverts indicate an adult of that sex; 3) adult male or adult female-type tertial coverts that differ from the surrounding immature-type coverts indicate the sex of some immature birds. 1 Wing Male Female Character Adult I 1 Immature I Immature 1 Adult Broad, smoothly rounded Appear narrow, and usually have ragged pointed tips Broad, smoothly rounded and flecked or vermiculated After molt: Similar to adult After molt: Similar to adult male female Figure 61. Immature female redhead SEPARATIONO F GREATER AND LESSER SCAUPS A white wing stripe normally extends onto the primaries on greater scaup but is confined to the secondaries on lesser scaup. There are exceptions. The stripes may not conform on a few male lesser scaup with unusually white wings or female greater scaup with unusually dark wings. Approximately 94 percent of scaup wings can be correctly assigned to species using wing length, provided the age-sex class of each wing is known. The following points will identify greater scaup in each age class: adult males > 213 mm., adult females > 206 mm., immature males > 209 mm and immature females > 203 mm. Wings in each age class that are shorter than these measurements are, in most cases, lesser scaup. Because the two species are similar, the following discussion pertains to both. Scaup differ from most other species of the same genus in that they rarely molt tertial coverts during the hunting season (October-January). Other characteristics useful for age-sex identification do not vary greatly, and once a person has learned to recognize them, wings of scaup are relatively easy to classify. Although charac-teristics that separate ages and sexes are similar for both species, they are not identical, and each is presented in a separate table. 11 wing 1 Male Character / Adult I Immature I Female 1 1 Immature I Adult I 1 I white near tios. tios oointed fleckina: tiosbointed. usuallv fleckina: tios oointed. usuallv without whiie fleckino 1 Middle and Black; all are well Black to dark brown with Black to dark brown, without Black to dark brown, plain or lesser vermiculated with white large white flecks to small white flecks; often ragged with very small white flecks coverts vermiculations recessed 118" and notched at tip concentrated near coverts' from coverts' edge; often edge; broadly rounded at tip raaaed and notched at ti0 Wing Male I Female Character Adult I Immature 1 Immalure 1 Adult / ~ c a ~ u l a r s ' Heavily vermiculated; appear Lightly venniculated or flecked with whlte; appear more dark than white more white than black I- After molt Similar to adult After molt: No obvious male chanpe 1 -- Tertials Black, flecked to Black, barely flecked to well Black to very dark brown, Black to very dark brown; vermiculated with white; tips vermiculated with white; tips without flecks of white; tips without flecks of white; tips pointed and often drooping usually frayed to sharp point usually frayed to sharp point usually bluntly polnted Middle and Black; all are well Black with large white flecks Black without white flecks; Tips broadly rounded and lesser vermiculated with white to small vermiculat~ons often ragged and notched at smooth; coverts black with coverts recessed 118" from coverts' tip small white flecks edge; often ragged and concentrated at tips or plain notched at ti^ Age Determination Ring-necked duck wings should be examined under ideal lighting, preferably daylight rather than artificial light. A high fraction of immatures replace tertial coverts during the hunting season with shiny black feathers that differ slightly from the very dark brown of the other wing coverts. It may be neces-sary to tilt a wing back and forth to see this differ-ence. If no difference is apparent and all tertial coverts are broadly rounded, the wing is from an adult. If, however, the tertial coverts tend to narrow toward their tips and/or appear frayed andlor notched, the wing is from an immature. Many of the middle and lesser coverts of immatures may also show notches. Sex Determination The sexes of approximately half of each age group can be identified from wing measurements provided all primary quills have hardened. Adults with a notch-length of 196 mm. or more are males and those with a notch-length of 188 mm. or less are females. Immatures with wings 194 mm. or longer are males while those with wings 184 mm. or shorter are females. There are a few exceptions to these measurements. There is too much overlap to permit accurate sex determination of adults with wings 189- 195 mm. long or immatures with wings 185-193 mm. long using measurements done. Adult male tertials are shiny, greenish black and bluntly pointed. The tertials of adult females are similar but slightly less shiny, greenish brown and broadly rounded. Males are usually flecked (often faintly) on the underwing over the radius and ulna. Females are rarely flecked in this area. All of these differences are slight and accurate sex determination is quite difficult. Because some immature males resemble immature females, sex determination of immatures is not completely reliable. Wing Male Female Character Adult 1 Immature I Immature 1 Adult Greater tertial Broadly rounded, dark Dark brown to blackish; usually narrow slightly to ragged tips Broadly rounded, dark ' coverts blackish After molt: Similar to adult After molt: Similar to adult brown male female 1 Underwing 1 White flecking present on small coverts near leading edge of Small coverts near leading edge brownish, with white edging, wing most pronounced near body 1 rarely with trace of white flecklng Notch-length 1 88% > 193 mm. .- f 8896 r 189 mm. 1 86% < 189 mm. 1 88% c 192 mm.
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Title | Species, age and sex identification of ducks using wing plumage pages 1 - 71 |
Contact | mailto:library@fws.gov |
Creator | Carney, Samuel M. |
Description | Carney1992_pages1-71.pdf |
FWS Resource Links | http://library.fws.gov |
Subject |
Document Birds |
Publisher | U.S. Fish and Wildlife Service |
Date of Original | 1992 |
Type | Text |
Format | |
Source | NCTC Conservation Library |
Rights | Public domain |
File Size | 11979941 Bytes |
Original Format | Document |
Length | 75 |
Full Resolution File Size | 11979941 Bytes |
Transcript | S DEPARTMENT OF THE INTERIOR S GEOLOGICAL SURVEY ATIONAL WETLANDS RESEARCH CENTER 30 CAJUNDOME BLVD 9FAYETTE LA 70506-31 52 BY Samuel M. Carney Washingon, D.C. 1992 U.S. Department of the Interior U.S. Fish and Wildlife Service P rocedures have been developed over the past 30 earsf for managing the take of waterfowl by hunters. Wings of ducks contributed voluntarily to the U. S. Fish and Wildlife Service by hunters are examined each year by experts. This procedure involves the determination of species, sex, and age of ducks through an examination of these detached wings. Many persons skilled at examining wings of ducks have retired in recent years. We became concerned that these skills might be lost to Jitture generations ij'not properly documented. Documenting such skills is dificult. Subtle dzferences in feather color andfeather texture are used to distinguish young ducks from adult ducks and males from females. Printed words and pictures have their limitations, but we hope this publication captures the technique and preserves it. /' Director U.S. Fish and Wildlife Service Introduction .......................................................... 1 Feather groups of the duck wing ............................ 2 Mallard .................................................................. 5 American black duck ........................................ 1 O Mottled duck ....................................................... 13 Gadwall .............................................................. 1 6 American wigeon ................................................. 19 Green-winged teal ............................................... 24 Blue-winged and cinnamon teals ......................... 28 Northern shoveler ................................................ 3 1 Northern pintail .................................................. 34 Wood duck .......................................................... 39 Harlequin duck ................................................... 42 Steller's eider ........................................................ 45 Separation of redhead and canvasback ................. 48 Redhead .............................................................. 49 Canvasback. ......................................................... 54 Separation of greater and lesser scaups ................. 59 Greater scaup ....................................................... 59 Lesser scaup ......................................................... 6 5 Ring-necked duck ............................................... 70 Separation of common and Barrow's goldeneyes ......................................... 75 Common goldeneye .........................................7..5. Barrow's goldeneye ...........................................-.8. 1 Separation of bufflehead and hooded merganser .........................................8..6 Bufflehead ........................................................8..7. Hooded merganser ............................................9..2 Separation of red-breasted and common mergansers .....................................9..6 Red-breasted merganser ......................................9. 6 Common merganser .......................................... 100 Separation of oldsquaw. black scoter. and surf scoter .............................................. 0 5 Oldsquaw .......................................................1..0.6 Black scoter ....................................................... 1 0 Surf scoter ......................................................... 114 White-winged scoter ........................................ 118 Common eider ............................................. 122 King eider .......................................................... 127 Ruddy duck ....................................................... 132 Fulvous whistling duck ...................................... 135 Black-bellied whistling duck .............................. 138 Appendix: Key to duck species .......................... 141 T his publication contains procedures used by the U.S. Fish and Wildlife Service to determine the species, sex and age composition of the harvest of North American ducks using detached wings contributed by hunters. Original studies of the use of duck wings for this purpose began in 1958 and were led by the author and A.D. Geis. Others con-tributing to these studies include R.L. Croft, E.M. Martin, A.N. Novara, L.D. Schroeder, M.G. Smart, and M.F. Sorensen. Major suppliers of known-age specimens include: W. Anderson, R.S. Billard, A.J.Erskine, D. Hall, A.S. Hawkins, C. Hoffpauir, L.R. Jahn, R. L. Jessen, F.B. Lee, J.J. Lynch, R.K. Martinson, D.I? Olson, R.I? Osbolt, C. Ritcey, R.N. Smith, H.E. Spencer, V.D. Stotts, and J. Takekawa. R.I. Smith edited the text, A.J. Godin prepared Figure 2, Larry Ketchum Photography took the pictures, and R.E. Cummins typed the manuscript. The Waterfowl Parts Survey became the means by which large samples of duck wings were obtained. This survey became national in scope in 1961. Collection of goose tails was added to the survey in 1962. Packages of envelopes are mailed to selected hunters who return wings from shot ducks and tail feathers from shot geese by mail to collection points throughout the United States where they are exam-ined to determine species, sex, and age. For readers who are interested in more detailed information on the development and testing of procedures described in the following pages, copies of a more technical report, which was prepared by the author, are available by writing to Waterfowl Harvest Surveys, U.S. Fish and Wildlife Service, 10800 Laurel-Bowie Road, Laurel, Maryland 20708-3600. That report, titled Observations on Sexing and Aging Ducks Using Wings, makes several points that must be understood by anyone who plans serious use of these techniques. The proce-dures presented here vary among species in the degree to which they accurately identify the age and sex of duck wings. Accuracy varies according to skills of the observer. Some individuals become highly skilled after examining large numbers of duck wings, while other individuals never develop the ability to detect subtle differences in feather texture and feather quality with high levels of profi-ciency. Levels of skill will decline if techniques are not practiced with regularity. Terminology related to age must be clarified. An immature wing possesses one or more characteristics known to be associated with ducks hatched in the most recent nesting season. Since the term imma-ture often refers to maturing processes not related to plumage, juvenalmight have been more appropri-ate. Nevertheless, after 30 years of use in this con-text, introducing another age designation would serve no purpose. All wings not possessing charac-teristics associated with the most recent hatch are classified as adult except those of male eiders which are identified as being from sub-adults. To determine the species, age, and sex of ducks from detached wings, a worker must be familiar with the various feather groups (Figure 1). The first step is to determine the species represented. A key to species is included as an appendix to this publica-tion. Usually, slight differences in feather shape, color, pattern, wear, or replacement are sufficient during the fall and winter to separate immatures from adults. Age determination is a step-by-step search for one or more traces of immature plumage. Wings on which no traces of immaturity can be found, or in some cases those that have positive adult characters, are considered to be from adults. During their first fall and winter immatures of many of the more common species of ducks molt certain wing-feather groups located near the body and replace them with adult-type feathers. Those feathers that are replaced include the tertials, greater tertial coverts, post humerals, and scapulars. Scapulars are of limited use in classifying wings because most hunters do not include scapulars on the wings they remove. Tertials, as defined here, are actually the more proximal secondaries, which are generally different in size, shape, and color from their more distal counterparts. These feathers are often sexually dimorphic and usually molt with adjacent body feathers. Post humerals are feathers attached to the humerus. They lie between the tertials and scapulars. They usually molt with adja-cent body feathers. Greater coverts are the first row continued on page 4 FEATHER GROUPS OF THE DUCK WING REMIGES: (Flight feathers) Alula: The feathered "thumb" of the bird wing Primaries: Flight feathers attached to the hand (manus) Secondaries: Flight feathers attached to the forearm (ulna) Tertials: Incorrect (morphologically) but used here to designate the more prox-imal secondaries which are generally different in size, shape, and color from their more distal counterparts, are often sexually dimorphic, and usually molt with adjacent body feathers. Post humerals: Feathers attached to the humerus. They lie between the tertials and scapulars, usually molting with adjacent body feathers. Scapulars: Feathers of the humeral (upper arm) feather tract. These lie on either side of the back and may partially cover a folded wing. They usually molt with adjacent body feathers. (Not shown) Axillars: Elongate feathers growing in the "armpit" region and closing the space between the spread wing and the body WING COVERTS: (Cover flight feathers) Greater coverts: The first row of feathers overlying the flight feathers, identified by the particular feathers they cover as primary, secondary, or tertial coverts. Tertial coverts: Those greater coverts that overlie the tertials. Designated separately here because they are sexually dimorphic in adults of several species and usually molt with the adjacent body feathers. Middle coverts: The next row of coverts. Lesser coverts: The next several rows of coverts. Marginal coverts: An indefinite number of rows anterior to the lesser coverts 2 PRIMARIES Figure 1. Feather groups of a typical dabbling duck wing of feathers overlying the flight feathers, identified by both fresh wings and wings with varying degrees of the particular feathers they cover as primary, sec- stiffness, as they are commonly received through the ondary, or tertial coverts. Greater tertial coverts are Waterfowl Parts Survey. To ensure uniformity all those greater coverts that overlie the tertials. They measurements are made using a standardized proce-are designated separately because they are sexually dure and measuring board (Figure 2). Measure-dimorphic in adults of several species and often ments are referred to as wing notch-length. molt with the adjacent body feathers. The degree to Tables have an advantage over keys in that the which these feathers are replaced is quite variable, color, shape, or texture of a particular group of I even among closely related species. Southern nest- feathers can be compared on one page across the ing duck species may initiate upperwing molts in four age and sex categories. Unlike keys, tables do the fall rather than in the spring. Fall wing molting not lead one directly to the answer. Despite this occurs among wood ducks, mottled ducks, and limitation, most people prefer tables to keys. whistling ducks. Such molting reduces the accuracy Therefore, tables are used in this publication to of immature wing identification in the fall and present information on individual species. Wing characteristics are not always listed in the same Male wings on most North American ducks are sequence for each species. They are listed in the slightly larger than those of females. For a few sequence in which they can be most efficiently used. species, this difference is large enough to permit A brief narrative, which identifies the most separation of the sexes using wing measurements. frequently used wing characters in a table, accompa- The procedure for measuring duck wings applies to nies most tables. Figure 2. Procedure for measuring wings 4 Place the heel of one hand over the end of measuring board and grasp wing in the area of the radius and ulna. Pull until wooden block seats tightly into the notch at the bend of the wing. Flatten the leading edge of the wing and note the length of the wing at the longest primary. Note: If primary quills are not completely grown, the measurernent is not val~d. Sex Determination The white bar anterior to the speculum extends onto the greater tertial coverts on all female wings but terminates at the proximal edge of the specu-lum on nearly all male wings. Approximately 2%-3 percent of males show some white edging on their tertial coverts. Adult males can be identified because the white is not continuous with that over the sec-ondaries. Immature males with white over the ter-tial coverts are difficult to tell from immature females. The white bar is the easiest sex character to use, because of its high degree of reliability and the fact that it is rarely lost when a wing is detached. Vermiculated scapulars are found only on males. Early in the hunting season (September and October) many males possess barred scapulars which are remnants of their summer plumage. Proximal underwing coverts are vermiculated or flecked on adult and most immature males. These feathers are barred on females and on a few imma-ture males. Age Determination of Males Immature tertials are often frayed and faded, usually narrow, and lack the pearly color of adult tertials. By late November immature tertials are replaced by first winter tertials, identical in appear-ance to adult feathers. At the same time, immature tertial coverts may be replaced by coverts which are broader, unfrayed, and similar to adult coverts and thus differ from adjacent immature coverts, which have not been molted. Many immature males have light edging on the inner webs of the most distal primary coverts. Adult males do not show this char-acter. Middle coverts of immatures are narrower and more trapezoidal than those of adults. Occasionally, these feathers on immature males are worn and/or have light edges. This type of edging does not occur on the middle coverts of adult males. Age Determination of Females Tertials that are frayed and/or faded are remnants of immature plumage and are found only on imma-ture ducks. Tertial coverts of immatures are often frayed, faded, and narrow, and the two most proxi-mal often lack the white of the speculum bar. As with males, greater tertial coverts of immatures may be replaced. Conspicuous light edging on the inner webs of the four most distal primary coverts is found only on immatures. Adults may have minute or no edging on these coverts. ~ i d d l ceo verts of immatures tend to be narrow and trapezoidal, while those of adults are broadly rounded. Scapulars 1 Venniculated o'barred or both / Barred + -. - I Wing Character I Proximal Flecked to vermiculated I Barred 1 I Male 1 Female Adult 1 Immature / Immature 1 Adult Tertials - - - - - - roiinded;.sotid gray to brown, no edgtljs Broad, pearly gray, no edging, rarely frayed or faded' May be growing in October Primary coverts Small, narrow, brownish, often frayed or faded near tips. Adul type feathers may be growing in December or later After mol: Similar to adul After mol: Similar to adul male female Broadty rounded; brown; ~suaHyw Sth light brown @iQlng; sometimes no SiigMfy ttiangukr to W o i d a l ; often frayed or faded; usually bmwn No edging Varies from pearly gray to brownish, often light edging; rarely frayed or faded Ma) have fiie'edging or no Edging varies from w9 Inner web of four most distal have llght edg~ngo r no edging conspicuous to absent edging I Inner web of four most distal wlthout edglng to conspicuous edging Innerweb of four most distal wlth llght edglng or no edging I m.; a IFigur-e 5. Adult hniale rnallrird ~ v i r hu necigid co\.ct-ts Figure 6. Immature male mallard with immature tertials Age Determination Sex Determination of Adults Because wings of the sexes are similar, it is easier Adult male tertials are more than 90 mm. long to determine the age of black ducks before attempt- from the edge of the longest tertial covert to the ing to identify their sex. Tertials small, narrow, and tertial tip, and acutely pointed with some pearly frayed andlor faded near their tips are remnants of color on the outer webs. Tertial coverts have broad immature plumage. Adult tertials are longer and edging which is pale brown. Middle and lesser wider and are not frayed or faded. Tertial coverts of coverts are broadly rounded and unfrayed. The the immature plumage tend to be narrow, some- notch-length of 94 percent of the adult male what trapezoidal, and frequently frayed andlor known-age specimens was greater than 28 1 mm. faded. Those of adults are wide, broadly rounded, Adult female tertials are less than 90 mm. from and rarely frayed or faded. During their first fall and the edge of the longest tertial covert to the tertial winter, a substantial fraction of the immatures may tip, and they are rather bluntly pointed. Pearly color replace both their immature tertials and tertial generally does not occur on the outer webs. Tertial coverts with adult-type feathers. Thus, it is impor- coverts have broad edging which is pale brown. tant to look closely at the middle coverts immedi- Middle and lesser coverts are broadly rounded and ately anterior to the tertial coverts for indications of unfrayed. The notch length of 94% of adult females the trapezoidal shape, duller color, and wear that was less than 281 mm. indicate immaturity. Primary coverts of many Immature tertials less than 88 mm. from the immatures have light edging on their inner webs. longest covert to the tertial tip are from females and This type of edging does not occur on adults. longer tertials are from males. 1 Wino 1 Male 1 Female 1 I Y"Y'YY'"' I Adult I Immature I Immature I Adult I 1 After molt: Similar to adult After molt: Similar to adult rounded'arc; not frayed of male female 1 faded 1 Greater 1 Four most distal do not have 1 Four most distal otten (but not always) have light edging on 1 Four most distal do not have I ure 11. Adult female black duck p n p y3qq spur a~naewwI' ~ ;a1rnS !j - - 1 Because mottled ducks are southern nesters, most females, but there is considerable overlap. Birds immatures have replaced both tertials and greater with three or more non-iridescent secondaries are tertial coverts at the time hunting occurs. Careful virtually all females, but birds with only one or two scrutiny, however, will usually reveal one or more non-iridescent secondaries may be of either sex. faded coverts in this area. These are remnants of the The greater tertial coverts of females tend to more immature plumage. Sexual identification is difficult. heavy edging than those of males, but the two types Wings of males are generally longer than those of grade together. Male I Female , Adult 1 Immature I Immature I Adult Similar to adult males; lack Similar to aduk females; I Often bluntly pointed; internal markings oflen internal markings often r mm.; lack internal . .- 4 Broadly rounded; edging Usually similar to those of Usually similar to those of Broadly rounded with heavy varies from broad to narrow adult males; often one or adult females; sometimes tan edging more narrow and frayed or one or more narrow and faded fraved or faded 1 Notch-length I 81% > 255 mm. 1 84% > 251 mm. 1 81% < 250 mm. 1 86% < 254 mm. 1 Sex Determination of Adults Greater, middle, and some lesser coverts of adult males are mostly either black or cinnamon. On adult females, black and cinnamon feathers are much restricted to the posterior three or four rows of coverts. Tertials of adult males are long, acutely pointed, silver-gay without edging or tipping. Those of adult females are much shorter, more bluntly pointed, and silver-brown with cream col-ored tips. Greater tertial coverts of adult males are part black and part gray, rarely with traces of white tipping. Those of adult females are similar but usu-ally well tipped with white. Marginal coverts of adult males are without edging but have arcs of narrow vermiculation. On adult females, these feathers are similar to the lesser coverts and ofcen have edging but may be either plain or with wide internal bars or arcs. Post humerals of adult females usually have cream edging at their tips. Those of other ages and sexes do not have this edging. Sex Determination of Immatures Tertials of both sexes are short, bluntly pointed, . . and ofieti frayed at their tips. They are quite similar to those of adult females. In late fall, they may be replaced by sexually dimorphic adult-type tertials. Greater tertial coverts of both sexes usually appear part black and part gray and are tipped with cream. They are narrower and more pointed than those of adults. Greater, middle, and sbme lesser coverts of immature males have some black andlor cinnamon in three or more rows. Immature females usually have little or no cinnamon color and black is ofien restricted to two rows of coverts. Some males show arcs of narrow vermiculation. Both sexes may show pale barring, which is usually wider on females. Notch-length measurements are usehl to identify immatures by sex, as long as the shafts of the pri-mary feathers have hardened. In 90 percent of the specimens measured, immature male notch-lengths were equal to or greater than 255 mm., while those of immature females were less than 255 mm. --- ------- Immature Immature 1 -- - Scapulars v o r b a n e d o r b a t h - - . -- - -- -- . I Barred Tertials Long, acutely pointed; Small; brownish; bluntly pointed; usually Shorter; bluntly pointed; silver-gray; without edging; ------ gray-brown with tips not hayed Or faded cream-colored tips which are not frayed or faded Greater tertial Bluntly pointed; outer webs Somewhat pointed; outer webs black; inner webs brown; Broadly rounded; usually coverts black; inner webs gray, often frayed and faded at tips with white tipping; outer 1 rarely with traces of white webs black or brown; inner tipping; not frayed or faded k - 7 After molt: Similar to adult After molt: Similar to adult fwaedbesd brown; not frayed or 1 Post 1 Broad, rounded, tips without 1 Narrow; pointed; tips without edaino 1 Broad rounded tips, usuallv 1 1 humerals edaing 1 1 lia- ht eda-ina" . I h- o.t z n a t h 7 5 e 2 6 2m m.. .. . . I 80% > 255 -mm. .. ( 80#s 255mm. I 9 ~ % c ~ r n m . 1 Adult males have a large white uppenving patch, long acutely pointed tertials with black outenvebs, and gray greater tertial coverts that are somewhat pointed and narrowly white-edged. Adult females have a sharply defined white edging on both greater and middle tertial coverts. Usually the sharp white edging on the middle and lesser coverts is sufficient to identify adult females, however, a few are so lightly colored as to resemble immature males. A carehl check of tertials and greater tertial coverts will serve to identify them. Immature males usually have small, brownish tertials and tertial coverts, but by November these are often replaced with adult male-type feathers. The middle and lesser coverts are gray-brown and indistinctly edged with a lighter gray. Immature females have small, brownish tertials and tertial coverts similar to those of immature males. By the middle of the fall hunting season these may be replaced with adult female-type feath-ers. The middle and lesser coverts are brownish with well-defined pale tan edges. Often the greater secondary coverts lack most of the black tipping common to wings of other ages and sexes and their outer webs are brownish gray to gray-white. Separation of American wigeon wings from Eurasian wigeon wings can be accomplished using the following procedures: American wigeon Axillars (if present) entirely white or flecked only at their tips. Undenving middle coverts all or nearly all white. Scapulars of males (if present) are heavily vermic-ulated with reddish brown. Eurasian wigeon Axillars (if present) heavily flecked with gray over their entire length. Underwing middle coverts heavily flecked with gray. Scapulars of males (if present) are heavily vermiculated with black and white. 1 Male KYaeter 1 Adult 1 Immature 1 Female I Immature I ~ d u ~ t I . -- Scapular~ I Vmiculated w barred or both -- I Barred A. Underwing Coverts gray, heavily flecked with white Coverts brownish; heavily edged with white to gray ( Occasionally as in females I Brown with light gray-brown edging; often frayed and faded Dark brown; edged with After molt: Similar to adult white to form a broadly I narrow white edging After molt: Similar to adult rounded arc male female GREEN-WINGEDT EAL If the scapulars are attached, the presence of one frayed. Adult female-type tertials have cream col-or more vermiculated feathers indicates that the ored edging. Adult male-type tertials are long, with-wing is from a male. If none of the scapulars are out edging, or with narrow edging. The presence of vermiculated, the wing could be that of either a adult-type tertials is not by itself an indication of male or female. age. Many geen-winged teal molt tertials during The stripe on the most distal tertial is the most the hunting season. For some of these, it is possible useful indicator of sex. On the wings of males, this to split incoming pin feathers to see whether the stripe is black and sharply delineated from the basic new feather is male or female in character. For feather color. On the wings of females, this stripe is wings with molted tertials, determination of sex is black to brown, but grades into the basic feather not possible. color. The sex of approximately half of the imma- Tertial coverts that are long and narrow, showing ture birds can be identified by measurements. fine, light edging, and a frayed fringe are remnants Wings 183 mm. or longer are from males and of immature plumage and positively identify the wings shorter than 175 mm. are from females, but wing as being immature. The tertial stripe may be sex cannot be determined from measurements used to separate the sexes. Adult female-type tertial between 176 mm. and 182 mm. A sample of wings coverts are broadly rounded with wide edging that from Adak Island, Alaska, green-winged teal (con- is cream or brown. Adult male-type tertial coverts sidered a different subspecies) averaged approxi- are a uniform gray, sometimes with a narrow bu@ mately 5 percent longer than wings collected in the edging, and tapering to a blunt point. Adult-type lower 48 states. The number of iridescent secon- tertial coverts are not a reliable indicator of age. daries differs between sexes. Wings having fewer Middle coverts of adult males are smooth and than four secondaries with completely iridescent without edging. Middle coverts of immatures are green outer webs are usually from females and rough and may show gray-appearing wear at their wings with five or more such secondaries are usually edges. Immatures may also possess light edging from males. Sex cannot be determined in this man- which is more pronounced on females, but may also ner from wings with four or four and a half irides- be present on males. Broad, rounded middle coverts cent secondaries. with wide edging are found only on adult females. Immature tertials are small, narrow, and rather Traces of immature plumage usually remain most delicate. The tips of these feathers are often badly obvious immediately anterior to the tertial coverts. Wing Male 1 Female Character Adult I Immature 1 Immature I Adult Tertials Uniform gray; tapering to a Small; narrow; rather delicate tips are often badly frayed; Buff edging including usually buff edging rounded tips; unfrayed; longitudinal stripe often poorly defined along inner edge; may be brown or black; stripe sometimes well defined; may be molting 1 1 ( male 1 female I I 1 Middle and Broadly rounded; no edging; Gray with wear around edges; appear ragged; somewhat Broadly rounded; usually lesser coverts gray that matches tertial narrow and trapezoidal; late in year contrast to a variable with wide light edgjng but coverts deoree with new freolaced) oreater coverts sometimes no eda~na Sex Determination The speculum of males is a bright iridescent green and that of females is a dull non-iridescent green. Rarely, a female may show a trace of irides-cence. The greater secondary coverts of males appear entirely white on their outer webs while those of females are heavily dark spotted and fre-quently appear more dark than light. A few males may show some spots in the greater coverts. Age Determination of Males Tertials of adults are greenish black, rarely frayed, andyery long and pointed. They are trimmed with a very narrow tan edging. Tertials of the immature plumage have wide edging which is tan and they are much shorter and more bluntly pointed than those of adults. They are brownish and often somewhat frayed and faded at the tips. Adult-type tertials occur on immatures and are similar to those of adult males but are usually still growing in December. Adult and replaced tertial coverts of immatures are similar, i.e. a dark brownish black without edging but usually showing a blue wash. Immature-type tertial coverts are brown, usually with pronounced edging and tan and usually not washed with blue. Age Determination of Females Tertials of adults are much more bluntly pointed than are the immature tertials. In addition, imma-ture tertials frequently become somewhat frayed at their tips. Tertial coverts of adults are usually straight sided to a rounded tip, whereas those of immature birds usually narrow slightly to a rounded tip that often shows traces of fading. Well propor-tioned white inverted "V's" generally indicate adults. -- -- --- , Immature I Adult Greater coverts Tertial Broadly rounded; dark coverts brown washed with blue; brown usually with usually without edging; not pronounced light brown frayed or faded I female The presence of white primary shafts separates shovelers from all other North American ducks. All birds having specula that are more than half gray or dull non-iridescent green are females. All males and a few adult females have iridescent green on more than half of their secondaries. Most females show cream edging on the lesser and middle coverts. This often covers all these feathers, particularly on adult females. Immature males may show a few cream colored edges on feathers near the alula. The tertials of immature males are brownish and much frayed, while those of adults and first winter immatures are greenish black and much longer. The tertial covcrts of the immature male are brownish black and often show a frayed fringe. Adult tertial coverts are blackish, often washed with blue. Both immature tertids and immature tertial coverts are usually present during October. Immature males generally have small dusky spots on their greater coverts, while adults do not. The immature tertials of females are similar to those of immature males. The tertials of adult females are wider, not frayed, and more heavily washed with white at the tips. Male I Female EFacIer 1 Adul, I Immature 1 Immature I Adult Smoothly rounded; dark Brown; often much frayed; with trace of light edging Brown; white edging forms brown to shiny black; After molt: Similar to adult After molt: Similar to adult smoothly rounded arc unfrayed male female White; dark bases normally White; often with small dark White; about 114 - 112 inch of dark base normally shows covered by middle coverts spots at tip; about 114 - 112 lnch of dark base normally shows 31 ' - Sex Determination one wing in 20 shows faint light edging, but the The speculum is at least partly iridesceRgreen general aspect of these is such that they can readily (rarely purple) on males and a dull non-iridescent be identified as adults. F bronze or rarely non-iridescent green on females. The tertial coverts of the immature plumage have The underwings of most males are flecked to ver- conspicuous edging which is a light yellowish miculated; those of a few males and all females are brown. Birds with such wings are immature males. barred. Male adult and first winter tertials are long Tertial coverts of the adult-type lack edging and are and gray and the innermost has a wide marginal found on both adult and immature wings. Tertials black stripe. The tertials of females are shorter and that lack the black edging are immature. It seems more brownish and lack the black stripe. Immature lilcely that wings with new tertials growing after tertials of both sexes are similar and somewhat like November are immatures. those of adult females. Vermiculated scapulars are found only on males, but unvermiculated sca~ulars Age Determination of Females may be present on both males and females. The middle coverts of the immature plumage are Tertials and scapulars are useful for identibing rather narrow and somewhat trapezoidal. Any bar-some males, but not for separating the sexes in all ring of these feathers is largely made up of triangu-cases. lar patches at the feather edge. The middle coverts of adults are broad and rounded and any barring is Age Determination of Males made up of variable shaped patches recessed from The most useful character for separating the ages the feather's edge. is the condition of the middle coverts. These have The tertial coverts of the immature plumage are light edging on immatures and often appear narrow, very frayed, often poorly edged and individual and frayed. Some of these feathers persist through- feathers are often long and narrow. The tertial out the hunting season. On adult males, these coverts of adult-type plumage are not very frayed, feathers are not frayed and generally plain. About are well rounded, and have conspicuous edging. Wing Male I Female Character 1 I Immature I Immature Adult I Underwing Gray; heavily flecked with white I Alternate brown and white bars I Tertials I Acutelv oointed: brownish I Brown: most distal with 1 Brown with oale lonaitudinal I Bluntlv oointed: dark brown: I I surrounding rachis I often frayed 1 frayed 1 brownish longitudinal stripe; I Middle and I Grav: without edaina: I Grav: liaht brown edaina at Brown with liaht edaina: 1 Brown with liaht edaina: edging; somewhat 1 from edging; rounded trapezoidal Immature f Adult Figure 42. Common middle covert markings of female northern pintail Sex Determination and extends farther distally than on wings of imma- The white trailing edge of the secondaries on tures. females is much wider on the outer webs than on the inner webs. On males, this white edge is Age Determination of Females approximately the same width on both webs. On Immature tertials are pale bronze with pointed, females, the first secondary proximal to these white- usually frayed, tips. Immature tertial coverts are edged feathers is washed with black on the outer narrow and yellow-geen. Most immatures replace web. Males have no black on this feather. both tertials and tertial coverts with purplish red adult-type feathers by late October. Usually, the Age Determination of Males blue iridescence is confined to two rows of coverts Immature tertials are pale bronze with pointed, and is restricted to the proximal half of the imma-usually frayed, tips. Immature tertial coverts are ture wing. Adult coverts are somewhat wider and narrow and yellow-green. Most immatures replace tend to lie more smoothly on the wing. Dark blue both tertials and tertial coverts with dark blue usually extends onto the third row of coverts and adult-type feathers by late October. As the imma- extends farther distally than on the immature ture middle and lesser coverts are replaced by the coverts. adult-type plumage, the presence of a few dark blue feathers among the duller coverts indicates immatu- Note: rity. Usually, the dark blue does not extend onto the By mid-fall, some immature wood ducks hatched third row of coverts and is much restricted to the in the southern United States appear to have area anterior to the ~roximahl alf of the secondaries. replaced all of their immature upper-wing coverts. Generally, coverts of adults are somewhat wider and Their wings cannot be distinguished from those of tend to lie more smoothly on the wing and the dark adults. blue usually extends on to the third row of coverts 1 Male I Female I , , blue-black; second most distal white-edged at itst ip A--fIt-e r molt: Similar to adult LAn-..lf.. ter molt: Similar to adult rounded tips 1 Other 1 Greater and middle coverts Greater coverts and middle Greater coverts often 1 Greater coverts usually 1 anteriorly over three or more confined to two rows of to two rows of cdverts three or more rows of rows of coverts coverts nearest secondaries anterior to secondaries coverts I pa A~J) IOU pue papunod h~qloo~us sd!~: 1 ~ 061 6u1peus' y3els SEPARATIONO F REDHEAD AND CANVASBACK Adult male canvasbacks have much more white secondary coverts and secondaries is greater on on the upper surface of their wings than any other wings of redheads of all age and sex groups than it is age-sex category of redhead or canvasback. All other on the wings of adult female or immature canvas-canvasbacks have much darker secondaries than backs of either sex. those of all redheads. Thus, the contrast between Immature greater coverts are narrower, squared, often frayed to a point over the tertials with an indistinct pale tip over the secondaries. During the hunting season, immature birds occasionally replace both their immature tertials and greater tertial coverts with adult-type feathers. For a given sex, these new feathers are indistinguishable from those of adults but differ markedly from other immature greater and middle coverts which are retained. With practice, redheads can be aged primarily by the appearance of their tertial coverts. Several combina-tions are possible: 1) immature-type tertial coverts always indicate an immature bird but sex is best determined from other coverts; 2) adult male or adult female-type coverts similar to the surrounding coverts indicate an adult of that sex; 3) adult male or adult female-type tertial coverts that differ from the surrounding immature-type coverts indicate the sex of some immature birds. 1 Wing Male Female Character Adult I 1 Immature I Immature 1 Adult Broad, smoothly rounded Appear narrow, and usually have ragged pointed tips Broad, smoothly rounded and flecked or vermiculated After molt: Similar to adult After molt: Similar to adult male female Figure 61. Immature female redhead SEPARATIONO F GREATER AND LESSER SCAUPS A white wing stripe normally extends onto the primaries on greater scaup but is confined to the secondaries on lesser scaup. There are exceptions. The stripes may not conform on a few male lesser scaup with unusually white wings or female greater scaup with unusually dark wings. Approximately 94 percent of scaup wings can be correctly assigned to species using wing length, provided the age-sex class of each wing is known. The following points will identify greater scaup in each age class: adult males > 213 mm., adult females > 206 mm., immature males > 209 mm and immature females > 203 mm. Wings in each age class that are shorter than these measurements are, in most cases, lesser scaup. Because the two species are similar, the following discussion pertains to both. Scaup differ from most other species of the same genus in that they rarely molt tertial coverts during the hunting season (October-January). Other characteristics useful for age-sex identification do not vary greatly, and once a person has learned to recognize them, wings of scaup are relatively easy to classify. Although charac-teristics that separate ages and sexes are similar for both species, they are not identical, and each is presented in a separate table. 11 wing 1 Male Character / Adult I Immature I Female 1 1 Immature I Adult I 1 I white near tios. tios oointed fleckina: tiosbointed. usuallv fleckina: tios oointed. usuallv without whiie fleckino 1 Middle and Black; all are well Black to dark brown with Black to dark brown, without Black to dark brown, plain or lesser vermiculated with white large white flecks to small white flecks; often ragged with very small white flecks coverts vermiculations recessed 118" and notched at tip concentrated near coverts' from coverts' edge; often edge; broadly rounded at tip raaaed and notched at ti0 Wing Male I Female Character Adult I Immature 1 Immalure 1 Adult / ~ c a ~ u l a r s ' Heavily vermiculated; appear Lightly venniculated or flecked with whlte; appear more dark than white more white than black I- After molt Similar to adult After molt: No obvious male chanpe 1 -- Tertials Black, flecked to Black, barely flecked to well Black to very dark brown, Black to very dark brown; vermiculated with white; tips vermiculated with white; tips without flecks of white; tips without flecks of white; tips pointed and often drooping usually frayed to sharp point usually frayed to sharp point usually bluntly polnted Middle and Black; all are well Black with large white flecks Black without white flecks; Tips broadly rounded and lesser vermiculated with white to small vermiculat~ons often ragged and notched at smooth; coverts black with coverts recessed 118" from coverts' tip small white flecks edge; often ragged and concentrated at tips or plain notched at ti^ Age Determination Ring-necked duck wings should be examined under ideal lighting, preferably daylight rather than artificial light. A high fraction of immatures replace tertial coverts during the hunting season with shiny black feathers that differ slightly from the very dark brown of the other wing coverts. It may be neces-sary to tilt a wing back and forth to see this differ-ence. If no difference is apparent and all tertial coverts are broadly rounded, the wing is from an adult. If, however, the tertial coverts tend to narrow toward their tips and/or appear frayed andlor notched, the wing is from an immature. Many of the middle and lesser coverts of immatures may also show notches. Sex Determination The sexes of approximately half of each age group can be identified from wing measurements provided all primary quills have hardened. Adults with a notch-length of 196 mm. or more are males and those with a notch-length of 188 mm. or less are females. Immatures with wings 194 mm. or longer are males while those with wings 184 mm. or shorter are females. There are a few exceptions to these measurements. There is too much overlap to permit accurate sex determination of adults with wings 189- 195 mm. long or immatures with wings 185-193 mm. long using measurements done. Adult male tertials are shiny, greenish black and bluntly pointed. The tertials of adult females are similar but slightly less shiny, greenish brown and broadly rounded. Males are usually flecked (often faintly) on the underwing over the radius and ulna. Females are rarely flecked in this area. All of these differences are slight and accurate sex determination is quite difficult. Because some immature males resemble immature females, sex determination of immatures is not completely reliable. Wing Male Female Character Adult 1 Immature I Immature 1 Adult Greater tertial Broadly rounded, dark Dark brown to blackish; usually narrow slightly to ragged tips Broadly rounded, dark ' coverts blackish After molt: Similar to adult After molt: Similar to adult brown male female 1 Underwing 1 White flecking present on small coverts near leading edge of Small coverts near leading edge brownish, with white edging, wing most pronounced near body 1 rarely with trace of white flecklng Notch-length 1 88% > 193 mm. .- f 8896 r 189 mm. 1 86% < 189 mm. 1 88% c 192 mm. |
Original Filename | Carney1992_pages1-71.pdf |
Date created | 2013-01-23 |
Date modified | 2013-03-06 |
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