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U. S. Fish and Wildlife Service American Woodcock Population Status, 2003 Suggested citation: Kelley, J.R., Jr. 2003. American woodcock population status, 2003. U.S. Fish and Wildlife Service, Laurel, Maryland. 20pp. All Division of Migratory Bird Management reports are available at our home page (http://migratorybirds.fws.gov). 1 AMERICAN WOODCOCK POPULATION STATUS, 2003 JAMES R. KELLEY, JR., U.S. Fish and Wildlife Service, Division of Migratory Bird Management, BHW Federal Building, 1 Federal Dr., Fort Snelling, Minnesota 55111-4056 Abstract: Singing-ground and Wing-collection surveys were conducted to assess the population status of the American woodcock (Scolopax minor). Singing-ground Survey data indicated that the number of displaying woodcock in the Eastern and Central Regions were unchanged from 2002 (P>0.1), although the point estimates of the trends were positive . Trends from the Singing-ground Survey during 1993-03 were –1.3 and –1.6% per year for the Eastern and Central regions, respectively (P<0.05). There were long-term (1968-03) declines (P<0.01) of 2.3% per year in the Eastern Region and 1.8% per year in the Central Region. The 2002 recruitment index for the Eastern Region (1.4 immatures per adult female) was the same as the 2001 index, but was 18% below the long-term regional average. The 2002 recruitment index for the Central Region (1.6 immatures per adult female) was 17% higher than the 2001 index (1.3 immatures per adult female), and was similar to the long-term regional average. The index of daily hunting success in the Eastern Region increased slightly from 1.8 woodcock per successful hunt in 2001 to 1.9 in 2002, but seasonal hunting success declined from 6.9 woodcock per successful hunter in 2001 to 6.6 in 2002. In the Central Region, the daily success index was 2.1 woodcock per successful hunt in 2001 and 2002; but seasonal hunting success increased from 10.0 woodcock per successful hunter in 2001 to 11.0 in 2002. The American woodcock is a popular game bird throughout eastern North America. The management objective of the U. S. Fish and Wildlife Service (FWS) is to increase populations of woodcock to levels consistent with the demands of consumptive and non-consumptive users (U. S. Fish and Wildlife Service 1990). Reliable annual population estimates, harvest estimates and information on recruitment and distribution are essential for comprehensive woodcock management. Unfortunately, this information is difficult and often impractical to obtain. Woodcock are difficult to find and count because of their cryptic coloration, small size, and preference for areas with dense vegetation. Up until the recent advent of the Harvest Information Program, a sampling frame for woodcock hunters had been lacking. Because of these difficulties, the Wing-collection Survey and the Singing-ground Survey were developed to provide indices of recruitment, hunting success and changes in abundance. This report summarizes the results of these surveys and presents an assessment of the population status of woodcock as of June 2003. The report is intended to assist managers in regulating the sport harvest of woodcock and to draw attention to areas where management actions are needed. METHODS Woodcock Management Units Woodcock are managed on the basis of 2 regions or populations, Eastern and Central, as recommended by Owen et al. (1977) (Fig. 1). Coon et al. (1977) reviewed the concept of management units for woodcock and recommended the current configuration over several alternatives. This configuration was biologically justified because analysis of band recovery data indicated that there was little crossover between the regions (Krohn et al. 1974, Martin et al. 1969). Furthermore, the regional boundaries conform to the boundary between the Atlantic and Mississippi Flyways. The results of the Wing-collection and Singing-ground surveys are reported by state or province, and region. Singing-ground Survey The Singing-ground Survey was developed to exploit the conspicuous courtship display of the male woodcock. Early studies demonstrated that counts of singing males provide indices to woodcock populations and could be used to monitor annual changes (Mendall and Aldous 1943, Goudy 1960, Duke 1966, and Whitcomb 1974). Before 1968, counts were conducted on non-randomly-located routes. Beginning in 1968, routes were relocated along lightly-traveled secondary roads in the center of randomly-chosen 10-minute blocks within each state and province in the central and northern portions of the woodcock’s breeding range (Fig. 1). Data collected prior to 1968 are not included in this report. The primary purpose of this report is to facilitate the prompt distribution of timely information. Results are preliminary and may change with the inclusion of additional data. Cover picture of incubating hen woodcock courtesy of Stephen Maxson, Minnesota Department of Natural Resources. 2 Each route was 3.6 miles (5.4 km) long and consisted of 10 listening points. The routes were surveyed shortly after sunset by an observer who drove to each of the 10 stops and recorded the number of woodcock heard peenting (the vocalization by displaying male woodcock on the ground). Acceptable dates for conducting the survey were assigned by latitude to coincide with peaks in courtship behavior of local woodcock. In most states, the peak of courtship activity (including local woodcock and woodcock still migrating) occurred earlier in the spring and local reproduction may have already been underway when the survey was conducted. However, it was necessary to conduct the survey during the designated survey dates in order to avoid counting migrating woodcock. Because adverse weather conditions may affect courtship behavior and/or the ability of observers to hear woodcock, surveys were only conducted when wind, precipitation, and temperature conditions were acceptable. The survey consists of about 1,500 routes. In order to avoid expending unnecessary manpower and funds, approximately one half of these routes are surveyed each year. The remaining routes are carried as “constant zeros.” Routes for which no woodcock are heard for 2 consecutive years enter this constant zero status and are not run for the next 5 years. If woodcock are heard on a constant zero route when it is next run, the route reverts to normal status and is run again each year. Data from constant zero routes are included in the analysis only for the years they were actually surveyed. Sauer and Bortner (1991) reviewed the implementation and analysis of the Singing-ground Survey in more detail. Trend Estimation.—Trends were estimated for each route by solving a set of estimating equations (Link and Sauer 1994). Observer data were used as covariables to adjust for differences in observers’ ability to hear woodcock. To estimate state and regional trends, a weighted average from individual routes was calculated for each area of interest as described by Geissler (1984). Regional estimates were weighted by state and provincial land areas. Variances associated with the state, provincial, and regional slope estimates were estimated using a bootstrap procedure (Efron 1982). Trend estimates were expressed as percent change per year and trend significance was assessed using normal-based confidence intervals. Short-term (2002-03), intermediate-term (1993-03) and long-term (1968-03) trends were evaluated. The reported sample sizes are the number of routes on which trend estimates are based. These numbers may be less than the actual number of routes surveyed for several reasons. The estimating equations approach requires at least 2 non-zero counts by the same observer for a route to be used. With the exception of the 2002-03 analysis, routes that did not meet this requirement during the interval of interest were not included in the sample size. For the 2002-03 analysis, a constant of 0.1 was added to counts of low-abundance routes to allow their use in the analysis. Each route should be surveyed during the peak time of singing activity. For editing purposes, “acceptable” times were between 22 and 58 minutes after sunset (or, between 15 and 51 minutes after sunset on overcast evenings). Due to observer error, some stops on some routes were surveyed before or after the peak times of singing activity. Earlier analysis revealed that routes with 8 or fewer acceptable stops tended to be biased low. Therefore, only route observations with at least 9 acceptable stops were included in the analysis. Routes for which data were received after 30 May 2003 were not included in this analysis but will be included in future trend estimates. Data for 2002 and 2003 were not received from Prince Edward Island. Therefore, short-term trends could not be estimated for the province; however, intermediate and long-term trends were estimated for 1993-2001 and 1968-2001, respectively. Annual indices.—Annual indices were calculated for the 2 regions and each state and province by finding the deviation between the observed count on each route and that predicted by the 1968-2003 regional or state/provincial trend estimate. These residuals were averaged by year and added to the fitted trend to produce annual indices of abundance for each region, state and province. Yearly variation in woodcock abundance was superimposed on the long-term fitted trends (see Sauer and Geissler 1990). Thus, the indices calculated with this method portray year-to-year variation around the predicted trend line, which can be useful for exploratory data analysis (e.g., observing periods of departure from the long-term trend). However, the indices should be viewed in a descriptive context. They are not used to CENTRAL EASTERN SURVEY COVERAGE BREEDING RANGE Fig. 1. Woodcock management regions, breeding range, and Singing-ground Survey coverage. 3 assess statistical significance and a change in the indices over a subset of years does not necessarily represent a significant change. Observed patterns must be verified using trend estimation methods to examine the period of interest (Sauer and Geissler 1990, Link and Sauer 1994). Wing-collection Survey The Wing-collection Survey was incorporated into a national webless migratory game bird wing-collection survey in 1997. Only data on woodcock will be presented in this report. As with the old survey, the primary objective of the Wing-collection Survey is to provide data on the reproductive success of woodcock. The survey also produces information on the chronology and distribution of the harvest and data on hunting success. The survey is administered as a cooperative effort between woodcock hunters, the FWS and state wildlife agencies. Participants in the 2002 survey included hunters who either: (1) participated in the 2001 survey; or (2) indicated on the 2001-02 Annual Questionnaire Survey of U. S. Waterfowl Hunters or Harvest Information Program Survey that they hunted woodcock. Wing-collection Survey participants were provided with prepaid mailing envelopes and asked to submit one wing from each woodcock they bagged. Hunters were asked to record the date of the hunt, and the state and county where the bird was shot. Hunters were not asked to submit envelopes for unsuccessful hunts. The age and sex of the birds were determined by examining plumage characteristics (Martin 1964, Sepik 1994) during the annual Woodcock Wingbee, a cooperative work session. Wings from the 2002-03 hunting season were accepted through 25 April 2003. The ratio of immature birds per adult female in the harvest provided an index to recruitment of young into the population. The 2002 recruitment indices were compared to long-term (1963-2001) averages. Annual indices were calculated as the average number of immatures per adult female in eac h state, weighted by the relative contribution of each state to the total number of wings received during 1963-2001 (to maintain comparability between years). Daily and seasonal bags of hunters who participated in the Wing-collection Survey in both 2001 and 2002 were used as indices of hunter success. These indices were weighted to compensate for changes in the proportion of the estimated woodcock harvest attributed to each state and adjusted to a base-year value (1969) for comparison with previous years (Clark 1970, 1972, 1973). Only data on successful hunts from prior years were used so that they would be comparable to data from the new survey. A successful hunt was defined as any envelope returned with complete information in which >1 woodcock wing was received. Harvest Information Program The Harvest Information Program (HIP) was cooperatively developed by the FWS and state wildlife agencies to provide reliable annual estimates of hunter activity and harvest for all migratory game birds (Elden et al. 2002). In the past, the annual FWS migratory bird harvest survey was based on a sampling frame that consisted solely of hunters who purchased a federal duck stamp. However, people that hunt only non-waterfowl species such as woodcock and doves are not required to purchase a duck stamp, and therefore were not included in that sampling frame. The HIP sampling frame consists of all migratory game bird hunters, thus it will provide more reliable estimates of woodcock hunter numbers and harvest than we have had in the past. Under this program, state wildlife agencies collect the name, address, and some additional information from each migratory bird hunter in their state, and send that information to the FWS. The FWS then selects random samples of those hunters and asks them to voluntarily provide detailed information about their hunting activity. For example, hunters selected for the woodcock harvest survey are asked to complete a daily diary about their woodcock hunting and harvest during the current year’s hunting season. Their responses are then used to develop nationwide woodcock harvest estimates. These estimates should be considered preliminary as refinements are still being made in the sampling frame and estimation techniques. RESULTS AND DISCUSSION Singing-ground Survey Trend Estimation.— The number of woodcock displaying during the 2003 Singing-ground Survey in the Eastern and Central Regions were not significantly different (P>0.1) from the 2002 levels, however the point estimate of the trends were positive (Table 1, Fig. 2). Trends for all states and provinces are reported in Table 1, but results based on fewer than 10 routes should be considered unreliable. Trends for the 1993-03 period were computed for 341 routes in the Eastern Region and 404 routes in the Central Region. Eastern and Central region breeding populations declined (P<0.05) 1.3 and 1.6% per year, respectively, during this period (Table 1). Long-term (1968-03) trends were estimated for 609 routes in the Eastern Region and 614 routes in the Central Region. There were long-term declines (P<0.10) in the breeding population throughout most states and CENTRAL EASTERN DECREASE (P<0.10) INCREASE (P<0.10) INSUFFICIENT SAMPLE SIZE DECREASE (NS) DECREASE (P<0.10) INSUFFICIENT SAMPLE SIZE INCREASE (NS) DECREASE (NS) Fig. 2. Short-term trends in the number of American woodcock heard on the Singing-ground Survey, 2002-2003. Fig. 3. Long-term trends in the number of American woodcock heard on the Singing-ground Survey, 1968-2003. CENTRAL EASTERN INCREASE (NS) 4 5 provinces in the Eastern and Central Regions (Table 1, Fig. 3). The long-term trend estimates were -2.3 and -1.8% per year (P<0.01) for the Eastern and Central regions, respectively. Annual Breeding Population Indices.—In the Eastern Region, the 2003 breeding population index of 1.78 singing-males per route was higher than the predicted value of 1.66 (Table 2, Fig. 4). The Central Region population index of 2.16 males per route was higher than the predicted value of 2.13. The major causes of these declines are thought to be degradation and loss of suitable habitat on both the breeding and wintering grounds, resulting from forest succession and various human uses (Dwyer et al. 1983, Owen et al. 1977, Straw et al. 1994). If current trends in land use practices persist, continued long-term population declines are likely. In an effort to halt such declines, the International Association of Fish and Wildlife Agencies has created a Woodcock Task Force to develop a woodcock conservation plan. Wing-collection Survey A total of 6,803 potential woodcock hunters in states with woodcock seasons were contacted and asked to participate in the 2002 Wing-collection Survey. Eighteen percent (Table 3) cooperated by sending in 9,002 woodcock wings (Table 4). Recruitment.—The 2002 recruitment index in the Eastern Region (1.4 immatures per adult female) was the same as the 2001 index, but was 18% below the long-term (1963-01) regional average of 1.7 immatures per adult female (Table 4, Fig 5). In the Central Region the 2002 recruitment index (1.6 immatures per adult female) was 17% higher than the 2001 index (1.3), and was similar to the long-term regional average. The preliminary 2002 recruitment index for eastern Canada (Ontario, New Brunswick, and Nova Scotia combined) was 2.5 immatures per adult female (Canadian Wildlife Service, unpublished data). Hunting Success.— The only change in Federal frameworks for woodcock hunting seasons in the U.S. during 2002-03 was moving the framework opening date in the Eastern Region from October 6 to October 1 (Appendix 1). The 2002 index of daily hunting success in the Eastern Region (1.9 woodcock per successful hunt) was slightly higher than the 2001 index of 1.8 (Table 5). The index of seasonal hunting success in the Eastern Region declined from 6.9 woodcock per successful hunter in 2001 to 6.6 in 2002. In the Central Region, the 2002 daily success index (2.1 woodcock per successful hunt) was similar to the 2001 index. Central Region hunters experienced an increase in the seasonal success index from 10.0 woodcock per successful hunter in 2001 to 11.0 woodcock per hunter in 2002. Base-year adjusted indices of daily and seasonal hunting success were below long-term averages in both regions (Figs. 6 and 7). NUMBER OF SINGING MALES PER ROUTE YEAR 1.0 1.5 2.0 2.5 3.0 3.5 4.0 4.5 68 71 74 77 80 83 86 89 92 95 98 01 1.0 1.5 2.0 2.5 3.0 3.5 4.0 4.5 EASTERN REGION CENTRAL REGION Fig. 4. Long-term trends (smooth line) and annual indices of the number of woodcock heard on the Singing-ground Survey, 1968-2003. 1.0 1.2 1.4 1.6 1.8 2.0 2.2 1.0 1.2 1.4 1.6 1.8 2.0 2.2 63 66 69 72 75 78 81 84 87 90 93 96 99 02 EASTERN REGION CENTRAL REGION YEAR ADJUSTED YOUNG PER ADULT FEMALE Fig. 5. Adjusted annual indices of recruitment (U.S.), 1963-2002. The dashed line is the 1963-2001 average. 6 Indices to seasonal hunting success indicate that the annual woodcock harvest has been declining among participants in the survey for over a decade. This is consistent with the results of the Annual Questionnaire Survey of U.S. Waterfowl Hunters (Martin 1979, and FWS unpublished data), which indicates that the woodcock harvest and the number of woodcock hunters have generally declined since the early 1980s (Fig. 8). These results should be interpreted cautiously because of the limitations of both of these surveys. A comprehensive critique of these limitations is beyond the scope of this report; interested readers should see Owen et al. (1977), Martin (1979), and Straw et al. (1994). Briefly, historic indices based on the Wing-collection Survey are potentially biased because of the non-random sampling procedure by which survey participants were selected. Because the Annual Questionnaire Survey of U. S. Waterfowl Hunters does not provide information on the woodcock harvest by non-waterfowl hunters, it does not provide an estimate of total harvest or the total number of hunters. Nevertheless, results from this survey should at least approximate trends in harvest and hunter participation. The 2001-02 estimates are the last ones generated from the Annual Questionnaire Survey, which has been replaced by HIP. Estimates of harvest and hunter numbers from the Annual Questionnaire Survey for 1964-2001 are presented in Tables 6 and 7 for historical purposes. 1.4 1.6 1.8 2.0 2.2 2.4 2.6 2.8 1.4 1.6 1.8 2.0 2.2 2.4 2.6 2.8 65 68 71 74 77 80 83 86 89 92 95 98 01 EASTERN REGION CENTRAL REGION ADJUSTED AVERAGE DAILY BAG YEAR Fig. 6. Base-year adjusted indices of daily hunting success in the U.S., 1965-2002. The base year is 1969; the dashed line is the 1965-2001 average. 1.4 1.6 1.8 2.0 2.2 2.4 2.6 2.8 1.4 1.6 1.8 2.0 2.2 2.4 2.6 2.8 65 68 71 74 77 80 83 86 89 92 95 98 01 EASTERN REGION CENTRAL REGION ADJUSTED AVERAGE DAILY BAG YEAR Fig. 7. Base-year adjusted indices of seasonal hunting success in the U.S., 1965-2002. The base year is 1969; the dashed line is the 1965-2001 average. 0 20 40 60 80 100 120 140 64 69 74 79 84 89 94 99 0 100 200 300 400 500 600 700 0 20 40 60 80 100 120 140 0 100 200 300 400 500 600 700 YEAR HARVEST (THOUSANDS) HUNTERS (THOUSANDS) HUNTERS HARVEST HUNTERS HARVEST EASTERN REGION CENTRAL REGION Fig. 8. U. S. harvest of American woodcock by duck stamp purchasers, and hunter numbers, 1964-2001 (Martin 1979, and FWS unpublished data, Division of Migratory Bird Management, Laurel, Maryland). 7 Harvest Information Program Estimates of active woodcock hunters, days afield, and woodcock harvest from the 2001-02 and 2002-03 HIP surveys are provided in Table 8. In the Eastern Region woodcock hunters spent approximately 161,200 days afield and harvested 84,800 birds during 2002-03. Woodcock hunters in the Central Region spent 476,800 days afield and harvested 235,100 birds during the 2002- 03 season. Although HIP provides statewide estimates of woodcock hunter numbers (Table 8), it is not possible to develop regional estimates, due to the occurrence of some hunters visiting more than one state to hunt. ACKNOWLEDGEMENTS Rebecca Rau (FWS) mailed and processed Singing-ground Survey forms, corresponded with cooperators, keypunched portions of data, participated in the wingbee, and continued development of the new web site that allowed cooperators to submit survey data electronically. T. Nguyen and H. Bellary (FWS) played vital roles in web site and database development. Personnel from the FWS, Biological Resources Division (BRD) of the U. S. Geological Survey, Canadian Wildlife Service (CWS), and many state and provincial agencies, and other individuals assisted in collecting the Singing-ground Survey data and processing wings at the Woodcock Wingbee. Special thanks to M. Bateman (CWS), G. Haas (FWS) and S. Kelly (FWS) for help in coordinating the Singing-ground Survey. Special appreciation is extended to A. Stewart, V. Tuovila and the Michigan Department of Natural Resources for hosting the 2003 wingbee. Individuals that participated in the wingbee were: D. Dessecker and A. Bump (Ruffed Grouse Society); F. Kimmel and M. Olinde (Louisiana Department of Wildlife and Fisheries); D. McAuley (BRD); W. Palmer (Pennsylvania Game Commission); A. Stewart and V. Tuovila (Michigan DNR); R. Stonebraker (Indiana DNR); E. Oppelt and J. Bruggink (Northern Michigan U.), M. Neal (U. of Michigan-Flint) and T. Edwards, C. Horton, J. Kelley, R. Rau, P. Stinson, R. Speer, K. Sturm and L. Wolff (FWS). Thanks to the 1,198 woodcock hunters that sent in wings. M. Bateman (CWS) provided preliminary estimates of woodcock recruitment for eastern Canada. The Harvest Surveys Section of the Division of Migratory Bird Management (FWS) mailed Wing-collection Survey materials, organized wing submissions, assisted with data management, and provided Harvest Information Program estimates of woodcock harvest (special thanks to P. Padding, M. Moore, E. Martin, and J. Bezek-Balcombe). E. Martin also provided the entire dataset on woodcock harvest and hunter numbers from the Annual Questionnaire Survey. B. H. Powell (BRD) developed the computer programs for administering the Wing-collection Survey. J. Sauer (BRD) developed computer programs for calculating trends and indices from Singing-ground Survey data. W. Kendall (BRD) performed the trend analyses and assisted with interpretation. R. Rau, W. Kendall, P. Padding, J. Sauer, and M. Otto reviewed a draft of parts or all of this report and provided helpful comments. Portions of this report were copied in whole or in part from previous woodcock status reports. LITERATURE CITED Clark, E. R. 1970. Woodcock status report, 1969. U. S. Fish and Wildl. Serv., Spec. Sci. Rep.—Wildl. 133. 35pp. _____. 1972. Woodcock status report, 1971. U. S. Fish and Wildl. Serv. Spec. Sci. Rep.—Wildl. 153. 47pp. _____. 1973. Woodcock status report, 1972. U. S. Fish and Wildl. Serv. Spec. Sci. Rep.—Wildl. 169. 50pp. Coon, R. A., T. J. Dwyer, and J. W. Artmann. 1977. Identification of harvest units for the American woodcock. Proc. American Woodcock Symp. 6:147-153. Duke, G. E. 1966. Reliability of censuses of singing male woodcock. J. Wildl. Manage. 30:697-707. Dwyer, T. J., D. G. McAuley, and E. L. Derleth. 1983. Woodcock singing-ground counts and habitat changes in the northeastern United States. J. Wildl. Manage. 47:772-779. Efron, B. 1982. The jackknife, the bootstrap and other resampling plans. Society for Industrial Applied Mathematics, Philadelphia, PA. 92pp. Elden, R.C., W.V. Bevill, P.I. Padding, J.E. Frampton, and D.L. Shroufe. 2002. Pages 7-16 in J.M. Ver Steeg and R.C. Elden, compilers. Harvest Information Program: Evaluation and recommendations. Int. Assoc. of Fish and Wildl. Agencies, Migratory Shore and Upland Game Bird Working Group, Ad Hoc Committee on HIP, Washington, D. C. 100pp. Geissler, P. H. 1984. Estimation of animal population trends and annual indices from a survey of call counts or other indicators. Proceedings American Statistical Assoc., Section on Survey Research Methods, 472-477. 8 Goudy, W. H. 1960. Factors affecting woodcock spring population indexes in southern Michigan. M. S. Thesis. Michigan State Univ., E. Lansing. 44pp. Krohn, W. B., F. W. Martin, and K. P. Burnham. 1974. Band recovery distribution and survival estimates of Maine woodcock. 8pp. In Proc. Fifth American Woodcock Workshop, Athens, GA. Link, W. A., and J. R. Sauer. 1994. Estimating equations estimates of trends. Bird Populations 2:23-32. Martin, E. M. 1979. Hunting and harvest trends for migratory game birds other than waterfowl: 1964- 76. U. S. Fish and Wildl. Serv. Spec. Sci. Rep. Wildl. 218. 37pp. Martin, F. W. 1964. Woodcock age and sex determination from wings. J. Wildl. Manage. 28:287-293. _____, S. O. Williams III, J. D. Newsom, and L. L. Glasgow. 1969. Analysis of records of Louisiana-banded woodcock. Proc. 3rd Annu. Conf. Southeastern Assoc. Game and Fish Comm. 23:85- 96. Mendall, H. L., and C. M. Aldous. 1943. The ecology and management of the American woodcock. Maine Coop. Wildl. Res. Unit. Univ. Maine, Orono. 201pp. Owen, R. B., Jr., J. M. Anderson, J. W. Artmann, E. R. Clark, T. G. Dilworth, L. E. Gregg, F. W. Martin, J. D. Newsom, and S. R. Pursglove, Jr. 1977. American woodcock (Philohela minor = Scolopax minor of Edwards 1974), Pages 149-186 in G. C. Sanderson, ed. Management of migratory shore and upland game birds in North America. Int. Assoc. of Fish and Wildl. Agencies, Washington, D. C. Sauer, J. R., and J. B. Bortner. 1991. Population trends from the American Woodcock Singing-ground Survey, 1970-88. J. Wildl. Mange. 55:300-312. _____, and P. H. Geissler. 1990. Estimation of annual indices from roadside surveys. Pages 58-62 in J. R. Sauer and S. Droege, eds. Survey designs and statistical methods for the estimation of avian population trends. U. S. Fish and Wildl. Serv., Biol. Rep. 90(1). 166pp. Sepik, G. F. 1994. A woodcock in the hand. Ruffed Grouse Society, Coraopolis, PA. 12pp. Straw, J. A., D. G. Krementz, M. W. Olinde, and G. F. Sepik. 1994. American woodcock. Pages 97-114 in T. C. Tacha and C. E. Braun, eds. Migratory Shore and Upland Game Bird Management in North America. Int. Assoc. of Fish and Wildl. Agencies, Washington, D. C. U. S. Fish and Wildlife Service. 1990. American woodcock management plan. U. S. Fish and Wildl. Serv., Washington, D. C. 11pp. Whitcomb, D. A. 1974. Characteristics of an insular woodcock population. Mich. Dept. Nat Resour., Wildl. Div. Rep. 2720. 78pp. 9 Table 1. Trends (% change per yeara and 90% confidence interval) in the number of American woodcock heard in the Singing-ground Survey as determined by the estimating equations technique (Link and Sauer 1994), 1968-2003. State, 2002-2003 1993-2003 1968-2003 Province No. of nc % change 90% CI n % change 90% CI n % change 90% CI CT 4 2 0.0 0.0 0.0 4 17.7 0.0 35.5 9 -10.0 ** d -16.5 -3.5 DE 2 0 2 -10.0 * -19.9 -0.1 2 4.2 -11.3 19.7 ME 44 29 23.1 * 1.0 45.1 53 -0.6 -2.0 0.8 64 -2.3 *** -3.1 -1.4 MD 6 2 -26.7 -110.8 57.4 6 -15.3 -43.0 12.5 21 -10.8 *** -17.2 -4.4 MA 7 6 -7.1 -43.8 29.6 9 5.2 ** 1.0 9.5 20 -4.1 ** -7.4 -0.7 NB 32 25 -2.1 -17.3 13.2 51 0.9 -1.5 3.3 62 -0.7 -1.9 0.5 NH 13 12 34.0 -8.6 76.7 13 0.7 -3.5 4.9 18 0.5 -2.2 3.3 NJ 7 2 28.7 -1.8 59.2 5 -4.1 -13.6 5.5 17 -10.1 *** -13.5 -6.8 NY 59 38 12.0 -14.7 38.6 70 -2.9 ** -4.9 -0.9 105 -2.8 *** -3.8 -1.8 NS 30 19 1.9 -23.9 27.7 38 1.1 -1.9 4.0 56 -0.3 -1.6 0.9 PA 34 13 -12.8 -41.9 16.3 26 -1.6 -7.2 4.1 56 -4.7 *** -6.9 -2.5 PEI 0 0 7 -2.8f -7.2 1.6 12 -1.4g -2.9 0.1 QUE 26 0 13 -2.9 ** -5.1 -0.7 54 -1.8 -4.9 1.2 RI 1 0 2 -15.7 *** -23.1 -8.2 VT 17 10 29.6 -11.2 70.3 18 0.7 -2.3 3.6 21 -1.5 -3.2 0.1 VA 26 4 17.0 -145.5 179.4 12 -10.7 * -20.2 -1.2 47 -10.7 *** -13.9 -7.4 WV 27 8 27.7 -58.1 113.6 14 -5.7 ** -9.8 -1.6 43 -2.7 *** -4.3 -1.0 Eastern 335 173 6.4 -4.9 17.7 341 -1.3 ** -2.4 -0.3 609 -2.3 *** -2.8 -1.8 IL 14 0 4 13.0 -17.2 43.2 23 23.4 -10.2 57.1 IN 17 2 0.0 0.0 0.0 7 -12.4 -28.3 3.6 38 -6.1 * -11.5 -0.7 MBe 18 9 31.7 -3.8 67.3 20 -4.0 * -7.8 -0.2 20 -3.5 ** -6.2 -0.9 MI 89 43 11.3 -6.1 28.6 109 -1.2 -2.6 0.1 143 -1.7 *** -2.5 -0.8 MN 72 46 5.5 -13.3 24.3 79 -0.4 -2.2 1.4 99 -1.1 ** -2.0 -0.3 OH 30 9 -9.9 -62.5 42.7 27 -10.6 ** -18.7 -2.6 55 -6.3 *** -10.0 -2.7 ON 34 0 87 -3.2 -7.9 1.5 136 -1.6 *** -2.3 -0.9 WI 69 41 7.9 -12.1 27.8 71 -1.7 -3.7 0.3 100 -1.9 *** -2.7 -1.0 Central 343 151 8.7 -1.7 19.2 404 -1.6 *** -2.5 -0.7 614 -1.8 *** -2.2 -1.3 Continent 635 324 7.7 * 0.7 14.8 745 -1.5 *** -2.2 -0.8 1223 -1.9 *** -2.3 -1.6 a Mean of weighted route trends within each state, province or region. To estimate the total percent change over several years, use: (100((% change/100)+1)y)-100 where y is the number of years. Note: extrapolating the estimated trend statistic (% change per year) over time (e.g., 30 years) may exaggerate the total change over the period. b Total number of routes surveyed in 2003 for which data were received by 30 May. c Number of comparable routes (2002 versus 2003) with at least 2 non-zero counts. d Indicates slope is significantly different from zero: * P<0.10, ** P<0.05. *** P <0.01; significance levels are approximate for states where n<10. e Manitoba began participating in the Singing-ground Survey in 1990. f Data were not received from PEI for the 2002 and 2003 surveys. Trend estimate is for the period 1993-2001. g Data were not received from PEI for the 2002 and 2003 surveys. Trend estimate is for the period 1968-2001. 10 1985 1.38 0.60 3.81 1.33 1.99 3.98 2.65 1.95 3.59 2.16 1.48 2.88 3.52 0.64 2.10 0.87 0.89 2.46 0.39 0.67 -- 4.89 3.66 1.47 5.02 3.03 2.98 2.53 1984 1.61 0.59 3.75 1.32 2.54 3.67 2.50 2.73 2.84 2.15 1.90 3.95 2.88 1.92 2.66 1.72 0.94 2.56 0.22 0.82 -- 4.68 3.06 1.70 4.88 3.28 2.78 2.59 1983 2.44 1.63 3.75 1.68 1.44 4.54 2.81 2.31 3.53 2.25 1.80 3.46 3.67 2.18 2.62 1.20 1.14 2.70 0.19 0.84 -- 4.25 3.44 1.83 4.64 3.00 2.87 2.65 1982 3.18 -- 2.92 2.52 1.93 4.33 2.40 1.99 3.06 1.80 1.59 2.18 2.96 3.19 1.79 1.55 1.10 2.44 0.13 0.80 -- 4.87 3.78 1.46 4.49 2.99 2.66 2.71 1981 2.57 -- 4.24 2.56 2.26 4.16 4.09 1.96 3.78 2.02 1.96 2.05 3.08 0.78 2.38 1.65 1.25 2.74 0.15 1.09 -- 4.61 4.19 2.03 5.92 3.05 3.17 2.78 1980 1.89 -- 3.87 3.00 2.20 4.12 3.91 2.55 4.16 2.18 1.98 2.72 3.96 1.35 2.67 1.68 0.91 2.77 0.10 1.05 -- 5.51 4.57 1.78 6.38 3.58 3.23 2.84 1979 2.16 0.41 4.40 2.43 3.06 4.61 3.22 4.12 3.61 2.29 2.17 3.67 3.62 1.35 3.00 1.98 1.10 2.91 0.08 1.43 -- 5.60 4.11 1.80 6.24 4.19 3.43 2.91 1978 2.18 0.48 4.03 3.08 2.80 4.15 3.14 2.41 3.20 2.82 1.91 2.96 3.57 0.78 3.11 1.81 0.76 2.63 0.07 1.15 -- 5.69 4.17 2.33 6.49 4.29 3.48 2.98 1977 3.60 0.51 4.37 2.90 2.40 5.80 3.12 4.16 4.04 2.44 2.40 3.70 2.92 -- 4.07 2.38 1.09 3.02 0.07 1.28 -- 5.38 4.16 2.93 6.00 4.07 3.50 3.05 1976 3.08 0.38 4.84 3.06 3.09 4.66 3.79 3.77 3.93 2.42 2.42 4.19 2.64 2.35 3.40 2.49 1.08 2.90 0.05 1.32 -- 5.88 4.20 2.57 5.51 3.75 3.42 3.12 1975 5.63 1.20 5.46 4.51 2.33 6.35 3.06 6.23 3.98 2.74 2.50 4.99 3.82 2.35 3.75 3.00 1.24 3.37 0.06 1.34 -- 6.39 4.17 2.37 5.74 3.91 3.60 3.19 1974 5.31 0.72 5.11 4.26 3.99 5.60 3.61 8.36 4.76 3.20 2.23 3.27 3.82 3.03 3.22 3.53 1.08 3.42 0.03 1.39 -- 6.35 4.80 3.16 6.53 4.03 3.66 3.27 1973 5.48 0.77 5.15 6.04 4.94 5.05 2.63 8.41 4.52 2.56 3.09 2.46 3.22 4.06 3.28 2.41 1.12 3.24 0.03 1.89 -- 5.47 4.13 2.46 6.09 3.94 3.52 3.34 1972 7.49 0.46 4.81 5.45 3.67 5.69 3.38 5.98 4.49 2.64 2.86 3.07 4.26 4.06 3.71 3.35 1.40 3.49 0.03 1.85 -- 5.33 3.63 2.97 6.82 3.87 3.60 3.42 1971 6.14 0.40 5.11 6.51 4.99 5.46 2.68 10.09 4.79 2.76 3.22 5.26 4.51 5.32 3.29 3.85 1.15 3.63 0.02 1.53 -- 5.63 4.27 3.55 6.15 4.05 3.66 3.50 1970 7.92 0.55 5.63 7.00 4.26 5.57 3.31 8.13 4.25 2.23 3.71 2.84 -- 2.82 4.30 4.83 1.19 3.72 0.02 1.98 -- 5.83 3.98 3.50 6.47 4.57 3.80 3.58 1969 7.94 0.48 5.36 7.89 3.63 5.25 2.87 6.50 5.54 2.62 3.38 3.81 -- 2.83 2.53 4.66 1.66 3.80 -- 2.08 -- 6.12 4.64 -- 6.81 4.22 3.90 3.67 Year 1968 --b 0.62 5.19 8.94 -- -- -- 7.52 5.03 3.61 3.67 -- -- -- -- -- 1.48 3.92 -- 2.40 -- 6.26 -- -- 6.22 4.28 3.91 3.76 Table 2. Breeding population indices for American woodcock from the Sing-ground Survey, 1968-2003. These indices are based on the 1968-2003 trend and should be used for exploratory data analysis only; observed patterns should be verified using trend estimation methods (Sauer and Geissler 1990). State, Province or Region Eastern Region CTa DEa ME MD MA NB NH NJ NY NS PA PEIa QUEa RIa VT VA WV Region Central Region IL IN MB MI MN OH ON WI Region Continent a Annual indices are unreliable due to small sample size. b Insufficient data. 11 2003 0.30 0.72 2.67 0.19 1.46 4.75 3.97 0.33 1.98 2.16 0.77 -- 2.54 0.02 2.08 0.16 0.62 1.78 6.96 0.29 1.93 3.53 2.95 0.47 4.18 2.32 2.16 1.66 2002 0.31 0.72 2.46 0.29 1.32 3.91 3.63 0.38 1.80 2.06 0.80 0.87 2.58 0.05 1.83 0.18 0.51 1.66 4.46 0.26 1.45 3.42 2.72 0.49 5.94 2.19 2.04 1.70 2001 0.33 0.67 2.58 0.63 1.28 4.89 3.41 0.58 2.02 2.56 0.76 3.05 2.29 -- 2.22 0.19 0.61 1.81 6.30 0.46 2.36 3.35 3.55 0.54 3.88 2.34 2.37 1.74 2000 0.89 0.93 2.99 0.35 1.38 4.54 3.22 0.59 1.97 2.75 0.61 2.84 2.43 -- 3.47 0.23 0.74 1.84 3.81 0.40 1.88 3.61 3.45 0.63 4.66 2.55 2.29 1.78 1999 1.32 0.42 3.06 0.34 2.05 4.96 4.69 0.84 2.19 2.32 0.92 2.42 2.95 -- 2.57 0.26 0.65 2.02 2.75 0.47 1.73 3.46 3.18 0.56 3.96 2.78 2.31 1.83 1998 0.62 1.42 2.41 0.24 1.35 3.99 3.81 0.67 2.24 2.32 1.17 2.86 2.35 -- 2.56 0.25 0.62 1.86 -- 0.73 1.80 4.30 3.22 0.72 3.99 2.30 2.44 1.87 1997 0.65 0.72 2.56 0.51 1.47 4.80 4.14 0.20 2.17 1.99 1.06 2.66 2.29 0.08 2.29 0.34 0.72 1.89 1.69 0.35 1.47 3.61 2.60 0.61 4.00 2.36 1.87 1.91 1996 0.74 0.72 2.31 0.44 1.37 3.93 3.76 0.97 2.19 2.61 0.99 3.06 1.19 -- 1.75 0.25 0.65 1.66 3.78 0.45 2.49 3.70 2.97 0.84 3.47 2.52 2.33 1.96 1995 0.81 -- 3.02 0.31 1.05 4.34 4.76 0.81 2.35 2.54 1.21 2.69 3.28 -- 2.32 0.29 1.04 2.17 1.21 0.53 2.77 3.86 3.29 0.80 4.75 2.40 2.43 2.00 1994 0.64 -- 2.84 0.52 1.44 5.14 2.42 0.34 2.26 2.07 0.65 2.31 2.75 -- 2.06 0.38 0.60 1.80 1.33 0.51 2.47 3.58 3.03 0.79 3.80 2.38 2.31 2.05 1993 0.51 -- 3.22 0.58 1.28 5.34 2.85 0.80 2.26 2.74 1.31 2.27 3.54 -- 2.04 0.49 0.69 2.16 1.27 0.59 3.43 3.95 3.47 0.95 4.38 2.55 2.63 2.10 1992 0.62 0.24 2.98 0.30 1.51 3.97 2.29 0.85 2.79 2.50 1.23 2.40 2.98 -- 1.93 0.43 0.78 2.07 1.06 0.55 2.61 3.93 3.26 0.92 4.85 2.59 2.51 2.15 1991 0.90 0.39 3.58 0.72 1.83 4.19 3.94 1.03 3.27 2.28 1.68 2.45 3.54 0.20 2.95 0.55 0.78 2.43 0.79 0.71 -- 5.50 3.84 1.04 5.05 3.25 2.98 2.20 1990 0.85 0.72 2.85 0.87 1.52 4.38 2.87 1.08 3.03 1.85 1.52 3.29 2.89 -- 3.01 0.57 0.85 2.21 0.52 0.69 -- 4.68 4.11 1.32 5.08 3.21 2.84 2.25 1989 0.98 -- 4.15 1.10 1.67 5.53 3.31 1.59 2.53 2.69 1.15 4.02 3.73 0.96 3.15 0.58 0.80 2.37 0.60 0.62 -- 4.79 3.57 1.01 5.39 3.29 2.82 2.30 1988 2.33 -- 4.06 0.97 2.12 4.26 3.19 1.67 3.27 2.47 1.60 4.23 2.51 0.96 3.39 0.66 0.79 2.39 0.49 0.61 -- 5.02 4.13 1.48 5.10 3.55 2.98 2.36 1987 0.92 -- 4.31 0.92 2.12 3.97 3.23 2.22 2.81 2.27 1.61 2.64 3.53 -- 2.86 0.93 0.99 2.53 0.48 0.65 -- 4.59 3.69 1.24 5.17 3.55 2.98 2.41 Year 1986 2.00 --b 3.94 1.17 2.04 3.33 4.51 1.90 3.06 2.53 1.70 3.78 3.39 0.64 2.66 0.90 0.86 2.50 0.32 0.90 -- 4.96 3.86 1.14 4.94 3.56 2.97 2.47 Table 2. Continued. State, Province or Region Eastern Region CTa DEa ME MD MA NB NH NJ NY NS PA PEIa QUEa RIa VT VA WV Region Central Region IL IN MB MI MN OH ON WI Region Continent a Annual indices are unreliable due to small sample size. b Insufficient data. 12 Table 3. Distribution of U.S. hunters contacted and hunters that submitted woodcock wings in the 2002-03 Wing-collection Survey. State of No. of hunters No. of hunters that Percent that residence contacted submitted wings submitted wings AL 32 0 0 AR 29 0 0 CT 207 21 10 DE 24 1 4 FL 98 0 0 GA 77 6 8 IL 161 4 2 IN 138 29 21 IA 73 6 8 KS 16 0 0 KY 32 2 6 LA 194 20 10 ME 403 119 30 MD 84 10 12 MA 421 70 17 MI 777 242 31 MN 535 75 14 MS 23 0 0 MO 145 13 9 NE 39 0 0 NH 224 77 34 NJ 180 37 21 NY 459 94 20 NC 104 5 5 ND 7 0 0 OH 190 20 11 OK 37 0 0 PA 495 61 12 RI 49 5 10 SC 78 10 13 TN 72 3 4 TX 71 3 4 VT 179 68 38 VA 167 18 11 WV 42 16 38 WI 941 163 17 Total 6,803 1,198 18 13 Table 4. Numbers of woodcock wings received from hunters, and indices of recruitment in the U.S. Recruitment indices for individual states were calculated as the ratio of immatures per adult female. The regional indices for 2002 were calculated as the average of the state values, adjusted for comparability with the 1963-2001 average. Recruitment indices were not calculated for states where the sample of wings was <125. State or Wings received Region of Total Adult females Immatures Recruitment index harvest 1963-01 2002 1963-01 2002 1963-01 2002 1963-01 2002 Eastern Region CT 13,149 89 2,909 19 8,066 57 2.8 DE 418 3 54 0 294 2 5.4 FL 660 0 150 0 410 0 2.7 GA 2,927 29 901 10 1,270 7 1.4 ME 73,296 969 21,567 301 36,667 486 1.7 1.6 MD 3,843 42 958 16 2,150 19 2.2 MA 19,253 428 5,820 142 9,550 182 1.6 1.3 NH 27,456 702 8,876 227 12,697 333 1.4 1.5 NJ 24,357 222 5,660 60 14,331 105 2.5 1.8 NY 51,328 748 16,975 251 23,890 317 1.4 1.3 NC 3,007 65 890 30 1,489 30 1.7 PA 27,589 351 8,705 109 12,777 146 1.5 1.3 RI 2,246 13 420 2 1,519 8 3.6 SC 2,312 105 712 25 1,102 44 1.5 VT 20,311 457 6,513 173 9,494 197 1.5 1.1 VA 3,948 121 942 53 2,274 44 2.4 WV 5,156 76 1,572 22 2,611 32 1.7 Region 281,256 4,420 83,624 1,440 140,591 1,872 1.7 1.4 Central Region AL 910 0 243 0 425 0 1.7 AR 515 0 165 0 207 0 1.3 IL 1,289 18 293 4 727 13 2.5 IN 6,819 138 1,717 37 3,802 61 2.2 1.6 IA 922 36 311 12 407 16 1.3 KS 44 0 9 0 22 0 a KY 1,010 23 238 6 524 15 2.2 LA 28,894 318 6,479 64 18,711 211 2.9 3.3 MI 100,620 2,239 32,536 677 50,320 1122 1.5 1.7 MN 28,604 538 9,708 224 12,840 183 1.3 0.8 MS 1,719 0 488 0 875 0 1.8 MO 2,703 84 668 22 1,360 45 2.0 NE 13 0 5 0 6 0 a OH 13,648 118 4,137 34 6,476 59 1.6 1.8 OK 170 2 38 0 89 2 2.3 TN 1,008 10 250 2 517 4 2.1 TX 986 1 261 1 501 0 1.9 WI 62,678 1,057 20,444 353 30,429 492 1.5 1.4 Region 252,552 4,582 77,990 1,436 128,238 2,478 1.6 1.6 14 Table 5. State and regional indices of daily and seasonal woodcock hunting success in the U.S. during 2001 and 2002. State and regional indices were calculated for states represented by >10 hunters that participated in the Wing-collection Survey both years. Regional indices were weighted as described by Clark (1970). State of No. of successful No. of successful hunts Woodcock bagged Woodcock per successful hunt Woodcock per season harvest hunters 2001 2002 2001 2002 2001 2002 2001 2002 Eastern Region CT 8 26 21 49 45 DE 1 2 1 4 3 GA 4 5 11 6 26 ME 84 391 363 875 795 2.2 2.2 10.4 9.5 MD 6 12 6 25 12 MA 33 132 152 247 275 1.9 1.8 7.5 8.3 NH 46 266 249 576 512 2.2 2.1 12.5 11.1 NJ 16 63 60 132 135 2.1 2.3 8.3 8.4 NY 60 278 290 507 559 1.8 1.9 8.5 9.3 NC 4 30 28 68 61 PA 39 143 129 319 272 2.2 2.1 8.2 7.0 RI 4 7 7 10 12 SC 9 27 43 60 99 VT 40 170 171 309 331 1.8 1.9 7.7 8.3 VA 7 45 45 104 86 2.2 2.2 12.6 10.1 WV 6 14 13 22 26 Region 367 1,611 1,589 3,313 3,249 1.8 1.9 6.9 6.6 Central Region IA 6 21 20 33 35 IL 3 6 9 10 15 IN 14 47 50 94 93 2.0 1.9 6.7 6.6 KY 2 15 12 31 23 LA 13 83 107 220 306 2.7 2.9 16.9 23.5 MI 186 1,076 955 2,224 1,932 2.1 2.0 12.0 10.4 MN 64 311 239 634 490 2.0 2.1 9.9 7.7 MO 5 22 27 50 60 OH 12 78 51 168 112 2.2 2.2 14.0 9.3 TN 2 14 5 25 7 TX 1 4 1 6 1 WI 120 545 406 1,111 840 2.0 2.1 9.3 7.0 Region 428 2,222 1,882 4,606 3,914 2.1 2.1 10.0 11.0 15 1982 10,981 1,637 419 4,355 4,437 18,376 7,788 12,951 6,856 15,450 33,513 8,959 33,329 1,283 4,459 6,042 2,770 2,874 176,479 2,874 4,802 12,707 7,131 4,015 952 3,864 112,186 60,222 39,761 8,926 4,076 862 246 12,273 904 98 4,739 5,893 43,688 330,219 1981 12,245 1,714 193 4,155 5,440 25,390 10,743 21,688 12,099 13,901 56,536 13,228 39,689 1,261 7,298 7,777 5,854 1,494 240,705 2,425 5,445 13,108 7,006 3,339 831 3,892 211,441 56,733 32,205 9,777 6,291 1,559 90 13,216 1,316 58 4,448 5,172 64,617 442,969 1980 15,217 2,929 184 5,251 6,691 30,132 8,329 20,109 12,780 27,108 46,196 6,973 23,740 1,186 7,503 6,185 4,691 3,230 228,433 5,132 6,389 11,346 10,965 6,904 785 2,883 327,751 71,999 52,627 21,742 8,516 1,055 326 14,931 1,265 161 2,822 16,900 77,072 641,571 1979 13,488 3,210 343 3,956 5,977 33,452 5,267 20,114 15,268 23,559 71,113 7,252 35,807 2,708 6,199 6,234 7,696 2,563 264,205 4,788 7,647 13,800 9,309 5,460 510 3,695 146,576 72,738 48,432 11,904 6,540 378 0 12,153 908 55 6,282 10,392 75,913 437,480 1978 15,905 4,349 399 5,667 6,747 36,279 11,261 23,949 19,860 30,933 48,286 9,897 37,391 2,213 8,926 8,567 6,628 3,418 280,675 4,514 6,081 19,533 8,396 8,662 935 4,963 214,793 98,260 53,865 11,042 12,929 1,203 292 12,112 1,384 0 9,375 6,689 96,926 571,954 1977 17,792 2,535 1,084 6,596 9,911 32,347 7,839 21,185 12,936 24,700 66,796 9,186 33,409 1,216 7,716 6,367 5,043 4,706 271,364 7,975 20,586 14,347 6,384 5,587 2,419 1,874 130,271 122,600 44,676 10,540 9,438 536 0 14,575 2,894 46 3,668 15,916 114,422 528,754 1976 20,134 4,165 191 8,881 8,623 35,440 9,883 28,492 16,399 37,937 59,342 8,936 43,550 2,624 7,288 16,443 9,421 3,868 321,617 5,796 8,162 10,481 10,583 5,025 1,115 4,260 121,740 128,568 18,019 17,651 8,733 540 150 14,891 2,779 135 4,082 12,207 125,453 500,370 1975 16,181 4,593 450 4,909 6,714 40,846 7,568 33,971 16,109 31,377 72,259 8,899 37,573 2,078 8,010 11,117 10,162 2,273 315,089 6,909 9,475 11,302 15,372 2,619 892 3,126 59,758 157,417 35,806 10,925 6,545 394 352 13,955 4,648 135 4,648 10,471 105,416 460,165 1974 13,547 3,772 500 5,402 10,999 48,123 8,530 31,648 16,754 42,308 83,600 14,617 45,170 3,457 9,363 9,234 5,392 2,345 354,761 3,532 6,811 11,480 12,163 4,584 2,297 2,330 67,492 114,106 25,290 11,082 6,145 520 378 16,906 2,117 0 5,724 5,516 101,895 400,368 1973 18,729 4,759 135 6,012 5,997 34,765 8,475 37,623 13,752 42,975 82,810 8,186 43,090 2,472 5,660 8,723 5,911 1,892 331,966 3,035 8,451 8,531 6,993 3,416 2,019 1,727 47,436 119,723 20,682 12,775 6,376 477 325 13,274 2,233 124 3,544 6,817 58,405 326,363 1972 12,953 2,057 975 5,439 8,186 35,762 4,382 34,958 12,098 23,906 77,935 9,970 49,599 2,238 9,280 7,061 4,461 4,305 305,565 6,721 8,139 10,509 8,797 2,945 1,131 790 134,592 95,512 26,589 12,997 2,880 489 825 10,511 1,696 82 4,257 5,428 82,265 417,155 1971 14,619 4,109 1,232 4,192 6,208 33,923 8,334 25,363 12,556 39,662 80,334 9,254 57,311 4,049 7,783 10,384 4,470 1,715 325,498 3,985 4,755 9,684 12,159 756 735 1,389 73,272 106,387 20,745 8,409 3,482 299 395 18,031 1,610 0 2,926 5,678 65,915 340,612 1970 23,430 3,626 360 6,868 5,579 33,171 9,781 32,197 9,520 28,030 74,873 9,181 54,754 2,585 9,050 11,459 4,120 1,454 320,038 5,424 3,723 7,589 6,293 1,085 1,073 788 95,777 77,671 20,721 9,534 4,294 1,333 74 13,136 836 93 1,727 4,353 90,768 346,292 1969 18,815 3,043 629 4,568 3,645 25,805 10,082 32,221 13,099 28,207 83,081 6,995 42,136 3,092 8,716 5,173 6,967 467 296,741 7,432 2,619 9,705 6,210 861 885 2,382 105,955 54,402 14,182 8,820 3,919 192 0 19,445 1,977 84 2,626 6,234 44,894 292,824 1968 15,772 3,798 330 5,799 6,226 38,617 9,127 29,330 15,483 24,622 76,528 9,612 30,917 2,086 8,741 4,553 4,342 1,213 287,076 4,114 3,193 2,083 3,884 483 629 692 75,271 42,244 7,948 6,337 1,529 33 348 15,345 535 32 2,602 2,804 27,794 197,900 1967 12,238 1,767 553 3,999 4,672 27,726 7,011 20,532 8,212 20,791 75,869 8,308 39,811 1,758 4,622 9,133 3,988 926 251,916 5,182 6,354 4,109 5,048 1,668 278 702 62,869 73,928 16,591 6,982 3,245 515 0 10,190 2,866 341 1,564 2,329 40,282 245,043 1966 13,137 2,210 747 3,453 2,594 25,350 3,103 25,056 9,490 14,094 49,459 9,042 18,236 1,584 3,000 7,178 3,260 413 191,406 5,458 2,684 4,133 4,145 222 268 667 53,842 68,661 9,952 4,556 2,392 56 441 9,776 386 90 1,097 4,972 36,190 209,988 1965 16,261 1,879 240 3,501 2,341 24,158 3,318 28,156 11,259 13,437 44,261 5,219 29,119 1,453 2,707 3,421 2,317 584 193,361 1,626 3,683 3,124 2,921 433 271 427 31,855 62,910 4,929 3,095 2,634 104 152 13,011 57 0 856 1,968 48,905 182,961 1964 11,928 1,804 91 2,126 2,893 26,115 3,025 19,696 6,488 11,108 38,625 6,790 19,340 541 8,552 4,233 2,648 186 166,189 2,224 3,952 4,015 4,761 1,563 547 737 71,744 90,662 10,795 7,390 2,646 253 44 16,905 327 100 488 8,115 58,934 286,202 Table 6. State and regional estimates of the number of American woodcock bagged (retrieved kill) by duck stamp buyers in the U.S., 1964-2001. Data from Martin (1979) and unpublished FWS administrative reports. Information was not available to reassign hunters from the District of Columbia to state of harvest. State or Region CT DE DC FL GA ME MD MA NH NJ NY NC PA RI SC VT VA WV Eastern Region AL AR IL IN IA KS KY LA MI MN MS MO NE ND OH OK SD TN TX WI Central Region 16 2001 3,748 392 121 4,924 1,990 14,087 1,973 5,511 3,792 4,623 5,593 4,226 11,866 187 4,626 3,446 3,280 923 75,311 124 231 3,256 4,974 948 442 3,008 7,026 35,290 16,044 955 3,646 823 238 6,280 1,318 117 1,341 2,727 14,253 103,041 2000 1,996 275 164 1,007 2,211 18,680 2,041 0 3,717 2,478 5,272 1,868 11,008 404 3,455 4,549 3,605 130 62,859 1,620 663 3,486 1,440 1,718 838 449 32,409 48,231 22,638 1,624 2,392 2,247 0 5,239 811 257 1,529 764 19,459 147,814 1999 5,306 0 214 787 1,503 15,028 2,099 5,818 2,668 4,286 8,886 4,403 8,020 619 5,061 3,711 10,025 2,221 80,654 80 4,053 3,071 2,520 895 3,303 979 23,066 61,992 23,456 326 1,684 0 0 6,273 1,264 0 4,293 2,576 22,709 162,540 1998 2,180 595 145 329 2,491 19,647 1,849 4,200 3,793 8,841 12,421 3,093 9,183 719 3,521 787 2,522 158 76,475 1,815 2,004 4,594 5,831 2,909 317 2,044 12,904 67,411 24,988 886 3,718 771 432 7,953 1,738 340 1,839 6,038 30,443 178,975 1997 2,718 477 216 975 898 15,479 2,034 5,233 3,037 4,131 10,735 2,962 9,550 203 2,521 2,688 3,234 0 67,091 1,117 1,299 4,152 3,022 2,478 792 412 12,645 45,417 29,865 5,627 2,483 2,185 0 6,487 585 21 9,116 6,002 30,714 164,419 1996 5,496 508 406 535 3,764 12,322 1,797 7,309 5,235 2,081 11,877 3,305 11,087 965 6,369 8,165 18,262 63 99,546 796 2,304 2,736 4,708 1,236 597 3,081 24,931 54,086 42,001 1,265 2,161 712 842 6,675 355 63 1,648 2,715 30,819 183,731 1995 6,209 641 137 187 1,989 18,227 3,253 12,216 5,023 4,338 11,955 2,803 7,918 538 5,244 4,274 2,943 389 88,282 599 274 5,023 4,978 1,864 883 1,708 67,428 70,329 31,409 2,708 2,043 162 255 4,721 630 41 2,813 2,985 28,706 229,559 1994 7,260 739 114 6,363 2,348 21,599 2,715 7,773 4,437 5,827 12,701 4,669 10,784 1,463 3,863 3,755 2,317 522 99,254 150 1,621 6,846 4,064 1,306 121 1,262 40,285 62,462 27,183 1,212 993 656 85 6,324 251 0 2,497 5,757 36,820 199,895 1993 4,630 317 149 1,422 2,843 14,551 3,198 8,378 4,158 7,882 10,199 2,789 13,476 481 3,032 2,384 1,898 35 81,822 356 248 5,849 2,914 1,172 225 1,265 44,096 54,058 20,260 889 1,962 153 569 7,950 524 0 1,009 2,109 32,076 177,684 1992 4,163 2,322 303 307 971 23,779 4,095 13,279 5,915 2,301 16,287 4,084 13,705 888 1,919 3,099 5,987 125 103,530 1,228 10,166 11,471 3,011 2,251 61 2,678 29,494 60,230 27,929 1,531 3,578 828 55 9,786 2,898 84 1,146 1,775 38,255 208,455 1991 5,174 1,281 120 741 2,513 20,550 3,506 13,751 6,616 7,821 12,371 4,750 11,719 2,061 1,759 4,411 4,343 650 104,259 651 1,780 9,804 2,431 1,972 612 1,932 37,289 66,833 35,917 994 3,349 303 701 8,518 716 1,521 1,763 3,775 36,968 217,829 1990 6,890 1,358 128 1,906 2,758 13,489 1,910 11,109 5,014 7,542 15,384 3,880 18,089 1,270 2,287 6,004 5,148 379 104,544 812 1,470 8,031 2,927 2,467 421 2,523 39,735 61,878 36,136 3,401 3,550 1,072 150 8,464 1,078 0 4,865 3,940 44,019 226,939 1989 5,884 3,230 0 2,405 2,863 16,093 6,574 13,083 5,808 10,900 20,041 5,159 15,220 491 4,518 5,098 4,684 854 122,904 1,886 1,220 8,935 6,685 5,457 616 3,343 66,615 50,856 45,396 2,065 6,279 1,124 156 15,165 2,268 219 3,996 5,434 42,965 270,680 1988 7,059 912 365 1,106 3,657 13,760 5,821 10,871 7,201 9,704 18,414 3,502 11,569 1,455 3,487 9,539 6,491 751 115,664 1,984 654 3,031 4,428 1,269 174 985 43,689 52,266 45,579 1,927 3,453 701 580 4,244 1,046 0 1,596 8,216 46,452 222,274 1987 7,374 1,694 370 4,270 2,203 27,404 9,887 15,793 6,567 10,885 27,073 5,373 21,354 2,530 6,569 7,198 6,256 409 163,209 3,298 5,682 7,672 8,194 3,897 2,108 1,287 123,958 70,062 71,966 2,891 7,818 481 0 5,866 1,097 580 3,506 9,498 61,375 391,236 1986 4,838 901 45 2,195 3,851 22,652 3,647 14,752 7,800 7,175 20,208 7,058 20,326 639 5,095 5,883 3,002 1,664 131,730 1,942 3,188 6,898 5,882 3,656 1,681 2,848 110,850 68,752 34,505 3,370 5,917 1,125 0 8,888 1,321 769 3,269 11,479 59,843 336,183 1985 6,319 849 65 2,608 2,988 17,314 7,221 17,139 7,420 8,804 17,945 7,299 16,214 1,988 2,094 4,439 7,112 578 128,394 2,108 6,636 7,285 4,921 3,888 1,260 1,991 78,532 56,885 28,678 6,388 4,087 364 754 8,719 2,043 0 3,235 7,922 42,360 268,056 1984 8,989 2,771 250 3,414 6,927 21,023 11,800 12,131 7,912 15,962 27,605 5,725 23,927 843 5,211 6,172 6,891 1,650 169,204 2,849 2,473 7,266 5,566 2,447 916 3,983 106,912 64,823 26,304 6,423 6,329 1,035 474 7,381 1,346 0 2,115 9,209 64,972 322,823 1983 9,378 1,210 0 5,661 5,231 19,155 6,812 11,937 4,701 11,391 27,789 5,859 27,340 1,915 7,137 4,651 2,783 3,293 156,244 2,408 2,584 8,047 4,865 2,731 1,164 3,412 196,201 55,615 35,346 4,839 4,133 3,257 264 12,941 1,408 275 2,479 8,510 44,112 394,591 Table 6. continued State or Region CT DE DC FL GA ME MD MA NH NJ NY NC PA RI SC VT VA WV Eastern Region AL AR IL IN IA KS KY LA MI MN MS MO NE ND OH OK SD TN TX WI Central Region 17 1982 3,749 778 76 828 1,509 5,058 2,037 6,169 2,967 3,993 13,536 2,676 14,972 578 1,770 2,152 1,434 564 64,846 798 1,720 3,711 2,102 1,463 512 872 19,233 14,406 11,791 1,424 1,839 435 227 4,490 338 87 1,239 1,824 16,851 85,362 1981 5,574 903 96 1,295 1,731 5,881 3,259 6,544 3,823 4,997 16,894 3,011 13,374 516 2,021 2,322 1,507 464 74,571 761 1,598 3,883 2,165 1,742 184 932 21,651 14,650 11,164 2,100 2,509 387 57 4,513 392 81 1,461 1,591 18,544 90,365 1980 5,173 905 73 1,366 2,091 5,920 2,117 5,609 3,526 4,983 15,150 2,470 13,522 397 2,569 2,197 1,447 513 70,030 1,067 1,976 4,340 2,485 2,443 419 706 27,508 17,000 17,098 2,544 2,780 325 137 4,770 444 73 1,176 3,068 22,111 112,470 1979 4,836 913 100 1,104 1,927 5,745 2,752 6,409 4,244 6,209 18,405 1,716 16,136 772 1,633 2,060 2,017 551 77,527 1,244 2,356 4,437 2,682 2,089 469 1,182 19,529 19,878 14,355 2,168 2,200 278 28 4,469 504 47 2,252 2,612 21,245 104,024 1978 5,614 1,439 199 1,382 1,746 6,910 3,115 6,026 4,448 8,227 17,330 2,815 19,591 763 2,603 2,460 1,986 501 87,155 1,406 1,642 6,729 2,794 4,110 510 1,750 24,041 20,271 16,925 2,683 3,288 564 182 3,678 776 31 2,378 2,556 20,489 116,803 1977 6,209 1,370 321 1,413 1,933 6,205 2,584 7,086 4,379 7,188 19,628 2,373 16,814 629 2,201 1,791 1,609 492 84,225 1,703 3,035 3,912 2,152 3,011 1,470 904 17,795 23,151 14,155 2,579 3,041 357 0 4,971 856 46 1,313 3,730 27,274 115,455 1976 5,987 1,675 93 2,042 1,428 6,397 3,675 7,824 4,761 8,337 17,345 2,857 18,700 990 1,987 2,782 2,132 457 89,469 1,459 2,666 4,108 2,717 2,619 796 1,249 16,463 30,685 7,857 2,864 2,387 270 150 5,623 898 67 1,274 3,176 35,332 122,660 1975 5,908 1,608 237 1,166 1,584 7,812 2,899 8,813 4,380 7,433 22,889 2,526 18,420 801 2,148 2,437 2,135 387 93,583 1,648 2,838 3,521 4,182 1,554 841 1,221 10,570 36,622 14,683 2,762 2,299 219 70 6,011 1,457 135 1,665 2,891 31,368 126,557 1974 5,390 1,423 204 1,753 2,084 7,306 3,084 9,521 4,253 8,459 25,384 3,007 20,221 1,075 2,628 2,376 1,721 425 100,314 1,191 2,348 4,544 3,666 2,138 1,364 775 10,063 33,455 10,233 2,120 1,837 271 168 6,052 628 37 1,751 1,976 30,420 115,037 1973 5,731 1,761 143 1,317 1,488 6,327 2,827 9,304 3,152 8,696 26,991 2,298 16,494 936 1,705 2,512 1,837 318 93,837 1,161 1,817 2,989 3,392 1,485 880 704 9,338 30,922 7,671 2,365 2,062 236 185 4,791 743 31 1,473 1,892 19,765 93,902 1972 5,006 1,138 195 1,265 1,872 5,503 2,013 9,187 3,388 6,925 24,284 2,337 17,662 729 2,136 1,731 1,733 688 87,792 1,585 2,693 4,205 3,227 1,374 638 429 16,039 28,312 8,621 2,486 1,414 163 214 4,524 716 41 1,416 2,016 24,667 104,780 1971 5,366 1,553 347 1,467 2,220 6,166 3,047 8,622 3,935 9,969 29,474 2,458 24,076 1,064 2,395 2,211 1,650 389 106,409 1,032 2,102 3,406 3,850 426 626 491 13,943 29,190 8,577 1,959 1,514 92 263 6,046 860 45 1,353 2,066 21,696 99,537 1970 7,286 1,403 180 2,961 1,558 5,798 3,252 10,148 2,973 7,728 25,789 2,748 22,030 794 2,480 1,859 1,611 482 101,080 1,449 1,117 2,483 2,784 452 1,893 354 17,636 29,894 9,360 2,195 2,053 172 37 5,268 611 46 624 1,753 27,885 108,066 1969 6,161 1,648 228 1,565 1,116 5,511 2,982 8,707 3,478 7,367 26,656 2,033 17,406 704 2,124 1,606 1,499 252 91,043 1,701 954 3,283 2,404 176 459 259 15,014 22,586 5,365 1,894 1,718 168 41 5,620 572 84 1,119 2,215 18,146 83,778 1968 4,450 1,163 157 1,576 1,429 5,522 2,551 7,884 2,613 5,815 21,977 2,138 12,069 483 2,449 1,578 1,187 314 75,355 1,571 912 1,347 1,444 170 463 293 9,733 17,711 3,756 848 762 33 194 3,561 376 32 571 875 12,182 56,834 1967 3,874 707 166 1,241 1,295 4,002 2,122 6,123 2,275 5,229 20,688 1,843 13,588 420 1,203 1,374 1,472 304 67,926 1,272 1,563 1,797 1,582 380 347 260 10,943 22,323 5,739 1,722 1,150 81 0 3,662 478 137 866 1,407 13,845 69,554 1966 3,388 829 197 1,331 820 4,115 1,462 6,787 2,128 4,525 16,712 2,169 7,688 492 1,165 1,608 843 197 56,450 1,424 639 1,454 1,387 121 245 351 11,112 20,480 2,932 1,337 934 56 110 2,958 192 45 676 1,578 11,730 59,761 1965 3,554 754 183 1,191 771 3,564 1,484 7,414 2,187 4,120 15,320 1,313 10,467 429 1,124 876 993 206 55,950 760 934 1,410 1,185 204 153 221 7,395 20,886 2,571 811 1,181 95 165 3,403 81 26 418 897 13,548 56,344 1964 3,129 706 91 758 812 2,980 1,407 5,548 1,594 3,462 12,699 1,735 7,317 382 1,398 1,017 1,086 93 46,214 828 1,394 1,328 1,480 570 364 224 10,327 23,631 4,536 1,422 1,255 207 88 5,357 265 33 298 1,799 18,257 73,663 Table 7. State and regional estimates of the number of duck stamp buyers that hunted American woodcock in the U.S., 1964-2001. Data from Martin (1979) and unpublished FWS administrative reports. Information was not available to reassign hunters from the District of Columbia to state of harvest. State or Region CT DE DC FL GA ME MD MA NH NJ NY NC PA RI SC VT VA WV Eastern Region AL AR IL IN IA KS KY LA MI MN MS MO NE ND OH OK SD TN TX WI Central Region 18 2001 1,353 245 53 352 1,045 3,168 1,382 1,510 1,083 1,046 3,493 1,209 5,277 187 1,292 920 719 212 24,546 400 135 1,655 1,353 772 97 891 2,130 12,961 6,923 917 1,304 396 160 3,221 325 95 903 1,720 7,322 43,680 2000 738 69 63 436 1,025 3,243 1,020 755 1,184 927 3,222 1,114 5,447 124 1,318 853 893 87 22,519 355 1,033 1,418 1,180 1,172 341 181 5,956 15,158 10,936 848 1,243 447 86 2,413 154 103 1,020 1,020 7,397 52,461 1999 1,305 0 50 346 586 3,791 855 2,033 956 1,282 4,193 1,529 5,885 221 1,477 941 1,564 119 27,131 323 1,637 1,417 1,304 721 930 839 5,617 17,539 8,197 177 998 196 111 2,503 520 0 1,145 734 8,670 53,578 1998 849 345 63 313 1,318 3,357 650 1,917 1,343 1,154 4,528 1,302 5,800 120 1,118 576 1,433 69 26,255 430 939 1,433 977 1,188 463 589 3,015 17,174 11,504 436 1,329 259 116 3,137 400 133 898 1,062 11,971 57,453 1997 1,470 286 74 598 758 2,903 607 1,772 1,031 1,269 4,439 1,192 5,753 174 1,045 950 1,305 142 25,767 374 542 1,846 1,010 1,244 275 311 2,816 16,789 11,455 639 1,408 400 75 3,600 250 76 1,016 1,453 11,389 56,968 1996 1,571 290 62 340 1,135 3,446 891 2,883 1,710 1,024 4,728 1,397 4,886 292 1,214 1,388 1,477 189 28,923 284 584 1,324 1,209 923 162 882 5,917 16,988 12,200 228 882 395 173 2,865 236 57 729 860 11,117 58,015 1995 1,929 401 48 189 837 3,009 1,161 2,992 1,511 1,298 4,837 1,017 5,670 269 950 1,300 1,004 158 28,581 331 241 1,831 1,207 1,060 273 909 5,654 18,503 11,745 574 774 185 225 2,617 378 85 899 318 11,115 58,924 1994 2,021 370 52 429 883 3,618 1,159 3,414 1,788 1,970 5,896 1,482 5,723 366 1,077 1,144 1,126 74 32,563 202 557 2,161 1,216 733 157 722 5,831 16,407 11,585 250 669 349 69 2,604 343 0 1,048 1,272 10,907 57,082 1993 1,916 226 60 463 1,217 3,718 1,125 3,617 1,652 2,378 5,122 1,172 6,464 262 1,153 840 944 71 32,397 250 113 2,118 1,424 489 160 569 7,988 16,547 7,933 330 1,051 282 269 3,080 405 0 536 969 11,060 55,573 1992 2,717 676 78 142 463 3,622 1,438 3,820 1,888 1,246 6,045 1,284 7,974 345 784 1,257 1,123 125 35,027 488 621 3,084 1,308 1,313 95 613 4,666 14,780 10,350 457 1,452 216 216 3,974 558 62 812 712 12,331 58,108 1991 3,016 620 45 549 838 4,016 1,494 4,195 1,666 1,793 6,244 1,855 7,701 420 951 1,485 1,427 104 38,464 337 914 3,575 1,125 1,292 140 704 5,981 16,445 11,735 417 962 177 201 3,075 333 129 717 1,288 12,787 62,334 1990 3,278 530 56 644 1,105 3,068 1,260 4,656 2,402 2,575 6,644 1,634 8,039 415 1,062 1,439 1,773 158 40,740 334 579 2,976 1,048 1,268 184 922 6,187 15,990 12,023 486 1,326 297 75 3,069 489 46 1,343 1,336 13,880 63,858 1989 2,958 392 33 551 1,036 3,043 1,365 4,816 2,029 2,672 7,901 1,752 7,550 404 1,457 1,299 1,540 197 40,995 730 573 2,913 1,499 1,506 338 910 8,478 14,296 14,212 676 1,796 260 262 4,189 620 125 837 1,519 14,353 70,092 1988 2,867 779 64 484 1,041 3,185 2,121 4,200 2,178 2,482 7,201 1,319 7,187 448 1,348 1,838 3,047 115 41,905 565 531 1,636 1,092 845 248 504 8,030 12,022 13,945 702 1,318 353 249 2,889 342 0 905 1,432 13,033 60,641 1987 3,687 731 74 1,075 1,311 4,195 2,842 5,117 2,433 2,743 9,716 1,809 10,055 567 1,912 1,977 2,317 203 52,764 605 1,638 3,078 1,722 1,957 812 746 14,792 14,563 14,076 828 2,252 507 41 3,350 502 137 1,276 2,264 17,084 82,230 1986 3,089 556 39 869 1,313 4,421 2,589 4,196 2,495 2,242 9,633 1,871 10,886 381 1,361 1,838 1,500 361 49,641 523 1,325 2,496 1,711 1,412 995 935 13,467 14,214 14,104 1,227 2,072 509 107 3,348 480 173 1,124 2,011 16,391 78,624 1985 3,390 438 49 658 1,312 4,196 3,147 4,733 2,380 3,274 9,418 1,885 9,546 478 1,266 1,629 2,168 213 50,182 731 1,404 2,533 1,349 1,622 600 698 13,907 13,888 10,524 1,116 2,141 374 177 3,879 668 27 1,010 1,502 14,874 73,024 1984 4,168 901 123 841 1,898 4,604 2,877 4,696 2,196 3,755 10,289 2,068 12,531 500 1,463 1,977 2,043 365 57,295 900 1,146 2,757 1,840 1,177 421 1,095 15,706 15,620 11,325 1,652 2,385 407 264 3,591 563 57 1,079 2,352 16,222 80,559 1983 4,385 691 0 1,057 1,901 4,459 2,512 4,129 2,460 4,411 10,515 2,066 12,816 619 2,054 1,828 1,011 543 57,459 675 1,094 2,938 1,529 1,546 415 858 20,526 12,642 10,414 1,394 1,565 737 175 4,188 647 132 765 1,931 15,052 79,223 Table 7. continued State or Region CT DE DC FL GA ME MD MA NH NJ NY NC PA RI SC VT VA WV Eastern Region AL AR IL IN IA KS KY LA MI MN MS MO NE ND OH OK SD TN TX WI Central Region 19 Table 8. Preliminary state and regional estimates of woodcock hunter numbers, days afield, and harvest from the 2001-02 and 2002-03 Harvest Information Program survey. Active woodcock hunters Days afield Harvest 2001-02 2002-03 2001-02 2002-03 2001-02 2002-03 Eastern region CT 1,800 ± 41% 1,600 ± 37% 7,500 ± 46% 9,200 ± 67% 3,600 ± 62% 4,500 ± 39% DE 400 ± 116% 500 ± 102% 5,200 ± 168% 900 ± 83% 200 ± 72% 500 ± 139% FL 2,400 ± 133% 1,000 ± 184% 14,600 ± 159% 2,000 ± 187% 9,500 ± 194% 100 ± 138% GA 3,600 ± 179% 2,600 ± 180% 40,500 ± 192% 5,600 ± 170% 11,000 ± 178% 600 ± 130% ME 11,900 ± 40% 5,000 ± 54% 64,900 ± 51% 18,500 ± 46% 48,100 ± 56% 18,600 ± 71% MD 700 ± 140% 600 ± 151% 1,500 ± 73% 1,100 ± 91% 1,700 ± 127% 600 ± 82% MA 1,200 ± 33% 1,100 ± 34% 5,700 ± 36% 6,000 ± 36% 2,500 ± 36% 3,500 ± 31% NH 2,000 ± 40% 1,500 ± 35% 9,900 ± 39% 7,500 ± 22% 6,700 ± 35% 5,600 ± 20% NJ 600 ± 66% 1,000 ± 69% 2,300 ± 23% 5,100 ± 86% 2,200 ± 30% 2,900 ± 57% NY 5,300 ± 37% 5,600 ± 36% 25,400 ± 41% 31,100 ± 47% 8,800 ± 55% 17,100 ± 62% NC 4,900 ± 154% 900 ± 67% 25,700 ± 147% 8,800 ± 104% 12,300 ± 126% 1,900 ± 132% PA 13,400 ± 45% 9,600 ± 44% 53,100 ± 52% 40,900 ± 57% 20,100 ± 52% 10,100 ± 40% RI 300 ± 88% 200 ± 82% 900 ± 101% 800 ± 73% 300 ± 63% 600 ± 83% SC 3,900 ± 92% 2,300 ± 129% 10,200 ± 107% 4,900 ± 122% 5,400 ± 171% 3,900 ± 163% VT 900 ± 39% 1,200 ± 45% 4,700 ± 36% 6,900 ± 55% 3,100 ± 28% 2,000 ± 31% VA 1,100 ± 128% 2,500 ± 86% 3,800 ± 108% 11,500 ± 96% 1,400 ± 30% 11,900 ± 176% WV 500 ± 82% 100 ± 21% 1,800 ± 100% 500 ± 30% 1,600 ± 73% 700 ± 42% Region na a na 277,800 ± 36% 161,200 ± 22% 138,100 ± 32% 84,800 ± 33% Central region AL 2,800 ± 109% 3,400 ± 93% 11,400 ± 146% 16,800 ± 95% 6,600 ± 191% 10,100 ± 109% AR 3,800 ± 131% 2,000 ± 172% 17,200 ± 166% 3,200 ± 113% 3,100 ± 132% 700 ± 112% IA 2,500 ± 78% 1,500 ± 103% 14,800 ± 113% 7,300 ± 134% 10,300 ± 128% 3,500 ± 130% IL 4,500 ± 81% 3,000 ± 90% 18,400 ± 82% 6,700 ± 86% 19,500 ± 112% 9,000 ± 111% IN 1,800 ± 106% 1,700 ± 113% 6,800 ± 118% 24,300 ± 172% 2,800 ± 96% 7,000 ± 160% KS 2,400 ± 110% 2,900 ± 96% 25,300 ± 113% 4,400 ± 111% 14,200 ± 138% 2,900 ± 137% KY 1,900 ± 174% 2,000 ± 126% 9,700 ± 171% 14,600 ± 150% 7,800 ± 171% 6,800 ± 141% LA 3,100 ± 139% 3,300 ± 148% 27,500 ± 155% 23,800 ± 166% 5,400 ± 59% 21,500 ± 138% MI 19,500 ± 23% 25,400 ± 18% 96,500 ± 24% 157,100 ± 37% 73,700 ± 33% 78,900 ± 26% MS 1,900 ± 133% 2,800 ± 187% 9,400 ± 154% 5,900 ± 179% 10,600 ± 129% 600 ± 64% MN 14,400 ± 49% 8,200 ± 66% 55,600 ± 47% 48,100 ± 94% 46,400 ± 71% 8,600 ± 31% MO 2,600 ± 101% 3,200 ± 125% 4,300 ± 89% 5,500 ± 115% 4,400 ± 114% 700 ± 40% NE < 50 ± 248% < 50 ± 155% 100 ± 73% 200 ± 72% 100 ± 99% 200 ± 74% OH 3,100 ± 135% 5,200 ± 108% 9,200 ± 93% 23,400 ± 137% 6,600 ± 87% 3,400 ± 43% OK < 50 ± 155% 2,500 ± 135% 200 ± 82% 6,300 ± 136% 100 ± 96% 2,600 ± 184% TN 200 ± 177% 4,200 ± 187% 700 ± 195% 4,800 ± 165% 700 ± 195% 8,000 ± 196% TX 10,400 ± 192% 28,700 ± 112% 12,800 ± 161% 67,000 ± 114% 5,300 ± 196% 38,300 ± 196% WI 14,800 ± 32% 17,100 ± 31% 68,700 ± 34% 57,500 ± 26% 33,700 ± 38% 32,000 ± 33% Region na na 388,600 ± 22% 476,800 ± 27% 251,400 ± 24% 235,100 ± 38% a Regional estimates of hunter numbers cannot be obtained due to the occurrence of individual hunters being registered in the Harvest Information Program in more than one state. 20 Appendix 1. History of framework dates, season lengths, and daily bag limits for hunting American woodcock in the U.S. portion of the Eastern and Central Regions, 1918-2002. Eastern Region Central Region Season Daily bag Season Daily bag Year (s) Outside dates length limit Year (s) Outside dates length limit 1918-26 Oct. 1 - Dec. 31 60 6 1918-26 Oct. 1 - Dec. 31 60 6 1927 Oct. 1 - Dec. 31 60 4 1927 Oct. 1 - Dec. 31 60 4 1928-39 Oct. 1 - Dec. 31 30 4 1928-39 Oct. 1 - Dec. 31 30 4 1940-47 Oct. 1 - Jan. 6 15 4 1940-47 Oct. 1 - Jan. 6 15 4 1948-52 Oct. 1 - Jan. 20 30 4 1948-52 Oct. 1 - Jan. 20 30 4 1953 Oct. 1 - Jan. 20 40 4 1953 Oct. 1 - Jan. 20 40 4 1954 Oct. 1 - Jan. 10 40 4 1954 Oct. 1 - Jan. 10 40 4 1955-57 Oct. 1 - Jan. 20 40 4 1955-57 Oct. 1 - Jan. 20 40 4 1958-60 Oct. 1 - Jan. 15 40 4 1958-60 Oct. 1 - Jan. 15 40 4 1961-62 Sep. 1 - Jan. 15 40 4 1961-62 Sep. 1 - Jan. 15 40 4 1963-64 Sep. 1 - Jan. 15 50 5 1963-64 Sep. 1 - Jan. 15 50 5 1965-66 Sep. 1 - Jan. 30 50 5 1965-66 Sep. 1 - Jan. 30 50 5 1967-69 Sep. 1 - Jan. 31 65 5 1967-69 Sep. 1 - Jan. 31 65 5 1970-71 Sep. 1 - Feb. 15 65 5 1970-71 Sep. 1 - Feb. 15 65 5 1972-81 Sep. 1 - Feb. 28 65 5 1972-90 Sep. 1 - Feb. 28 65 5 1982 Oct. 5 - Feb. 28 65 5 1991-96 Sep. 1 - Jan. 31 65 5 1983-84 Oct. 1 - Feb. 28 65 5 1997 *Sep. 20 - Jan. 31 45 3 1985-96 Oct. 1 - Jan. 31 45 3 1998 *Sep. 19 - Jan. 31 45 3 1997-01 Oct. 6 - Jan. 31 30 3 1999 *Sep. 25 - Jan. 31 45 3 2002 Oct. 1 - Jan. 31 30 3 2000 *Sep. 23 - Jan. 31 45 3 2001 *Sep. 22 - Jan. 31 45 3 2002 *Sep. 21 - Jan. 31 45 3 * Saturday nearest September 22.
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Title | American woodcock population status, 2003 |
Contact | mailto:library@fws.gov |
Description | Am_woodcock_population03.pdf |
FWS Resource Links | http://library.fws.gov |
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Document Birds |
Publisher | U.S. Fish and Wildlife Service |
Date of Original | 2003 |
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Transcript | U. S. Fish and Wildlife Service American Woodcock Population Status, 2003 Suggested citation: Kelley, J.R., Jr. 2003. American woodcock population status, 2003. U.S. Fish and Wildlife Service, Laurel, Maryland. 20pp. All Division of Migratory Bird Management reports are available at our home page (http://migratorybirds.fws.gov). 1 AMERICAN WOODCOCK POPULATION STATUS, 2003 JAMES R. KELLEY, JR., U.S. Fish and Wildlife Service, Division of Migratory Bird Management, BHW Federal Building, 1 Federal Dr., Fort Snelling, Minnesota 55111-4056 Abstract: Singing-ground and Wing-collection surveys were conducted to assess the population status of the American woodcock (Scolopax minor). Singing-ground Survey data indicated that the number of displaying woodcock in the Eastern and Central Regions were unchanged from 2002 (P>0.1), although the point estimates of the trends were positive . Trends from the Singing-ground Survey during 1993-03 were –1.3 and –1.6% per year for the Eastern and Central regions, respectively (P<0.05). There were long-term (1968-03) declines (P<0.01) of 2.3% per year in the Eastern Region and 1.8% per year in the Central Region. The 2002 recruitment index for the Eastern Region (1.4 immatures per adult female) was the same as the 2001 index, but was 18% below the long-term regional average. The 2002 recruitment index for the Central Region (1.6 immatures per adult female) was 17% higher than the 2001 index (1.3 immatures per adult female), and was similar to the long-term regional average. The index of daily hunting success in the Eastern Region increased slightly from 1.8 woodcock per successful hunt in 2001 to 1.9 in 2002, but seasonal hunting success declined from 6.9 woodcock per successful hunter in 2001 to 6.6 in 2002. In the Central Region, the daily success index was 2.1 woodcock per successful hunt in 2001 and 2002; but seasonal hunting success increased from 10.0 woodcock per successful hunter in 2001 to 11.0 in 2002. The American woodcock is a popular game bird throughout eastern North America. The management objective of the U. S. Fish and Wildlife Service (FWS) is to increase populations of woodcock to levels consistent with the demands of consumptive and non-consumptive users (U. S. Fish and Wildlife Service 1990). Reliable annual population estimates, harvest estimates and information on recruitment and distribution are essential for comprehensive woodcock management. Unfortunately, this information is difficult and often impractical to obtain. Woodcock are difficult to find and count because of their cryptic coloration, small size, and preference for areas with dense vegetation. Up until the recent advent of the Harvest Information Program, a sampling frame for woodcock hunters had been lacking. Because of these difficulties, the Wing-collection Survey and the Singing-ground Survey were developed to provide indices of recruitment, hunting success and changes in abundance. This report summarizes the results of these surveys and presents an assessment of the population status of woodcock as of June 2003. The report is intended to assist managers in regulating the sport harvest of woodcock and to draw attention to areas where management actions are needed. METHODS Woodcock Management Units Woodcock are managed on the basis of 2 regions or populations, Eastern and Central, as recommended by Owen et al. (1977) (Fig. 1). Coon et al. (1977) reviewed the concept of management units for woodcock and recommended the current configuration over several alternatives. This configuration was biologically justified because analysis of band recovery data indicated that there was little crossover between the regions (Krohn et al. 1974, Martin et al. 1969). Furthermore, the regional boundaries conform to the boundary between the Atlantic and Mississippi Flyways. The results of the Wing-collection and Singing-ground surveys are reported by state or province, and region. Singing-ground Survey The Singing-ground Survey was developed to exploit the conspicuous courtship display of the male woodcock. Early studies demonstrated that counts of singing males provide indices to woodcock populations and could be used to monitor annual changes (Mendall and Aldous 1943, Goudy 1960, Duke 1966, and Whitcomb 1974). Before 1968, counts were conducted on non-randomly-located routes. Beginning in 1968, routes were relocated along lightly-traveled secondary roads in the center of randomly-chosen 10-minute blocks within each state and province in the central and northern portions of the woodcock’s breeding range (Fig. 1). Data collected prior to 1968 are not included in this report. The primary purpose of this report is to facilitate the prompt distribution of timely information. Results are preliminary and may change with the inclusion of additional data. Cover picture of incubating hen woodcock courtesy of Stephen Maxson, Minnesota Department of Natural Resources. 2 Each route was 3.6 miles (5.4 km) long and consisted of 10 listening points. The routes were surveyed shortly after sunset by an observer who drove to each of the 10 stops and recorded the number of woodcock heard peenting (the vocalization by displaying male woodcock on the ground). Acceptable dates for conducting the survey were assigned by latitude to coincide with peaks in courtship behavior of local woodcock. In most states, the peak of courtship activity (including local woodcock and woodcock still migrating) occurred earlier in the spring and local reproduction may have already been underway when the survey was conducted. However, it was necessary to conduct the survey during the designated survey dates in order to avoid counting migrating woodcock. Because adverse weather conditions may affect courtship behavior and/or the ability of observers to hear woodcock, surveys were only conducted when wind, precipitation, and temperature conditions were acceptable. The survey consists of about 1,500 routes. In order to avoid expending unnecessary manpower and funds, approximately one half of these routes are surveyed each year. The remaining routes are carried as “constant zeros.” Routes for which no woodcock are heard for 2 consecutive years enter this constant zero status and are not run for the next 5 years. If woodcock are heard on a constant zero route when it is next run, the route reverts to normal status and is run again each year. Data from constant zero routes are included in the analysis only for the years they were actually surveyed. Sauer and Bortner (1991) reviewed the implementation and analysis of the Singing-ground Survey in more detail. Trend Estimation.—Trends were estimated for each route by solving a set of estimating equations (Link and Sauer 1994). Observer data were used as covariables to adjust for differences in observers’ ability to hear woodcock. To estimate state and regional trends, a weighted average from individual routes was calculated for each area of interest as described by Geissler (1984). Regional estimates were weighted by state and provincial land areas. Variances associated with the state, provincial, and regional slope estimates were estimated using a bootstrap procedure (Efron 1982). Trend estimates were expressed as percent change per year and trend significance was assessed using normal-based confidence intervals. Short-term (2002-03), intermediate-term (1993-03) and long-term (1968-03) trends were evaluated. The reported sample sizes are the number of routes on which trend estimates are based. These numbers may be less than the actual number of routes surveyed for several reasons. The estimating equations approach requires at least 2 non-zero counts by the same observer for a route to be used. With the exception of the 2002-03 analysis, routes that did not meet this requirement during the interval of interest were not included in the sample size. For the 2002-03 analysis, a constant of 0.1 was added to counts of low-abundance routes to allow their use in the analysis. Each route should be surveyed during the peak time of singing activity. For editing purposes, “acceptable” times were between 22 and 58 minutes after sunset (or, between 15 and 51 minutes after sunset on overcast evenings). Due to observer error, some stops on some routes were surveyed before or after the peak times of singing activity. Earlier analysis revealed that routes with 8 or fewer acceptable stops tended to be biased low. Therefore, only route observations with at least 9 acceptable stops were included in the analysis. Routes for which data were received after 30 May 2003 were not included in this analysis but will be included in future trend estimates. Data for 2002 and 2003 were not received from Prince Edward Island. Therefore, short-term trends could not be estimated for the province; however, intermediate and long-term trends were estimated for 1993-2001 and 1968-2001, respectively. Annual indices.—Annual indices were calculated for the 2 regions and each state and province by finding the deviation between the observed count on each route and that predicted by the 1968-2003 regional or state/provincial trend estimate. These residuals were averaged by year and added to the fitted trend to produce annual indices of abundance for each region, state and province. Yearly variation in woodcock abundance was superimposed on the long-term fitted trends (see Sauer and Geissler 1990). Thus, the indices calculated with this method portray year-to-year variation around the predicted trend line, which can be useful for exploratory data analysis (e.g., observing periods of departure from the long-term trend). However, the indices should be viewed in a descriptive context. They are not used to CENTRAL EASTERN SURVEY COVERAGE BREEDING RANGE Fig. 1. Woodcock management regions, breeding range, and Singing-ground Survey coverage. 3 assess statistical significance and a change in the indices over a subset of years does not necessarily represent a significant change. Observed patterns must be verified using trend estimation methods to examine the period of interest (Sauer and Geissler 1990, Link and Sauer 1994). Wing-collection Survey The Wing-collection Survey was incorporated into a national webless migratory game bird wing-collection survey in 1997. Only data on woodcock will be presented in this report. As with the old survey, the primary objective of the Wing-collection Survey is to provide data on the reproductive success of woodcock. The survey also produces information on the chronology and distribution of the harvest and data on hunting success. The survey is administered as a cooperative effort between woodcock hunters, the FWS and state wildlife agencies. Participants in the 2002 survey included hunters who either: (1) participated in the 2001 survey; or (2) indicated on the 2001-02 Annual Questionnaire Survey of U. S. Waterfowl Hunters or Harvest Information Program Survey that they hunted woodcock. Wing-collection Survey participants were provided with prepaid mailing envelopes and asked to submit one wing from each woodcock they bagged. Hunters were asked to record the date of the hunt, and the state and county where the bird was shot. Hunters were not asked to submit envelopes for unsuccessful hunts. The age and sex of the birds were determined by examining plumage characteristics (Martin 1964, Sepik 1994) during the annual Woodcock Wingbee, a cooperative work session. Wings from the 2002-03 hunting season were accepted through 25 April 2003. The ratio of immature birds per adult female in the harvest provided an index to recruitment of young into the population. The 2002 recruitment indices were compared to long-term (1963-2001) averages. Annual indices were calculated as the average number of immatures per adult female in eac h state, weighted by the relative contribution of each state to the total number of wings received during 1963-2001 (to maintain comparability between years). Daily and seasonal bags of hunters who participated in the Wing-collection Survey in both 2001 and 2002 were used as indices of hunter success. These indices were weighted to compensate for changes in the proportion of the estimated woodcock harvest attributed to each state and adjusted to a base-year value (1969) for comparison with previous years (Clark 1970, 1972, 1973). Only data on successful hunts from prior years were used so that they would be comparable to data from the new survey. A successful hunt was defined as any envelope returned with complete information in which >1 woodcock wing was received. Harvest Information Program The Harvest Information Program (HIP) was cooperatively developed by the FWS and state wildlife agencies to provide reliable annual estimates of hunter activity and harvest for all migratory game birds (Elden et al. 2002). In the past, the annual FWS migratory bird harvest survey was based on a sampling frame that consisted solely of hunters who purchased a federal duck stamp. However, people that hunt only non-waterfowl species such as woodcock and doves are not required to purchase a duck stamp, and therefore were not included in that sampling frame. The HIP sampling frame consists of all migratory game bird hunters, thus it will provide more reliable estimates of woodcock hunter numbers and harvest than we have had in the past. Under this program, state wildlife agencies collect the name, address, and some additional information from each migratory bird hunter in their state, and send that information to the FWS. The FWS then selects random samples of those hunters and asks them to voluntarily provide detailed information about their hunting activity. For example, hunters selected for the woodcock harvest survey are asked to complete a daily diary about their woodcock hunting and harvest during the current year’s hunting season. Their responses are then used to develop nationwide woodcock harvest estimates. These estimates should be considered preliminary as refinements are still being made in the sampling frame and estimation techniques. RESULTS AND DISCUSSION Singing-ground Survey Trend Estimation.— The number of woodcock displaying during the 2003 Singing-ground Survey in the Eastern and Central Regions were not significantly different (P>0.1) from the 2002 levels, however the point estimate of the trends were positive (Table 1, Fig. 2). Trends for all states and provinces are reported in Table 1, but results based on fewer than 10 routes should be considered unreliable. Trends for the 1993-03 period were computed for 341 routes in the Eastern Region and 404 routes in the Central Region. Eastern and Central region breeding populations declined (P<0.05) 1.3 and 1.6% per year, respectively, during this period (Table 1). Long-term (1968-03) trends were estimated for 609 routes in the Eastern Region and 614 routes in the Central Region. There were long-term declines (P<0.10) in the breeding population throughout most states and CENTRAL EASTERN DECREASE (P<0.10) INCREASE (P<0.10) INSUFFICIENT SAMPLE SIZE DECREASE (NS) DECREASE (P<0.10) INSUFFICIENT SAMPLE SIZE INCREASE (NS) DECREASE (NS) Fig. 2. Short-term trends in the number of American woodcock heard on the Singing-ground Survey, 2002-2003. Fig. 3. Long-term trends in the number of American woodcock heard on the Singing-ground Survey, 1968-2003. CENTRAL EASTERN INCREASE (NS) 4 5 provinces in the Eastern and Central Regions (Table 1, Fig. 3). The long-term trend estimates were -2.3 and -1.8% per year (P<0.01) for the Eastern and Central regions, respectively. Annual Breeding Population Indices.—In the Eastern Region, the 2003 breeding population index of 1.78 singing-males per route was higher than the predicted value of 1.66 (Table 2, Fig. 4). The Central Region population index of 2.16 males per route was higher than the predicted value of 2.13. The major causes of these declines are thought to be degradation and loss of suitable habitat on both the breeding and wintering grounds, resulting from forest succession and various human uses (Dwyer et al. 1983, Owen et al. 1977, Straw et al. 1994). If current trends in land use practices persist, continued long-term population declines are likely. In an effort to halt such declines, the International Association of Fish and Wildlife Agencies has created a Woodcock Task Force to develop a woodcock conservation plan. Wing-collection Survey A total of 6,803 potential woodcock hunters in states with woodcock seasons were contacted and asked to participate in the 2002 Wing-collection Survey. Eighteen percent (Table 3) cooperated by sending in 9,002 woodcock wings (Table 4). Recruitment.—The 2002 recruitment index in the Eastern Region (1.4 immatures per adult female) was the same as the 2001 index, but was 18% below the long-term (1963-01) regional average of 1.7 immatures per adult female (Table 4, Fig 5). In the Central Region the 2002 recruitment index (1.6 immatures per adult female) was 17% higher than the 2001 index (1.3), and was similar to the long-term regional average. The preliminary 2002 recruitment index for eastern Canada (Ontario, New Brunswick, and Nova Scotia combined) was 2.5 immatures per adult female (Canadian Wildlife Service, unpublished data). Hunting Success.— The only change in Federal frameworks for woodcock hunting seasons in the U.S. during 2002-03 was moving the framework opening date in the Eastern Region from October 6 to October 1 (Appendix 1). The 2002 index of daily hunting success in the Eastern Region (1.9 woodcock per successful hunt) was slightly higher than the 2001 index of 1.8 (Table 5). The index of seasonal hunting success in the Eastern Region declined from 6.9 woodcock per successful hunter in 2001 to 6.6 in 2002. In the Central Region, the 2002 daily success index (2.1 woodcock per successful hunt) was similar to the 2001 index. Central Region hunters experienced an increase in the seasonal success index from 10.0 woodcock per successful hunter in 2001 to 11.0 woodcock per hunter in 2002. Base-year adjusted indices of daily and seasonal hunting success were below long-term averages in both regions (Figs. 6 and 7). NUMBER OF SINGING MALES PER ROUTE YEAR 1.0 1.5 2.0 2.5 3.0 3.5 4.0 4.5 68 71 74 77 80 83 86 89 92 95 98 01 1.0 1.5 2.0 2.5 3.0 3.5 4.0 4.5 EASTERN REGION CENTRAL REGION Fig. 4. Long-term trends (smooth line) and annual indices of the number of woodcock heard on the Singing-ground Survey, 1968-2003. 1.0 1.2 1.4 1.6 1.8 2.0 2.2 1.0 1.2 1.4 1.6 1.8 2.0 2.2 63 66 69 72 75 78 81 84 87 90 93 96 99 02 EASTERN REGION CENTRAL REGION YEAR ADJUSTED YOUNG PER ADULT FEMALE Fig. 5. Adjusted annual indices of recruitment (U.S.), 1963-2002. The dashed line is the 1963-2001 average. 6 Indices to seasonal hunting success indicate that the annual woodcock harvest has been declining among participants in the survey for over a decade. This is consistent with the results of the Annual Questionnaire Survey of U.S. Waterfowl Hunters (Martin 1979, and FWS unpublished data), which indicates that the woodcock harvest and the number of woodcock hunters have generally declined since the early 1980s (Fig. 8). These results should be interpreted cautiously because of the limitations of both of these surveys. A comprehensive critique of these limitations is beyond the scope of this report; interested readers should see Owen et al. (1977), Martin (1979), and Straw et al. (1994). Briefly, historic indices based on the Wing-collection Survey are potentially biased because of the non-random sampling procedure by which survey participants were selected. Because the Annual Questionnaire Survey of U. S. Waterfowl Hunters does not provide information on the woodcock harvest by non-waterfowl hunters, it does not provide an estimate of total harvest or the total number of hunters. Nevertheless, results from this survey should at least approximate trends in harvest and hunter participation. The 2001-02 estimates are the last ones generated from the Annual Questionnaire Survey, which has been replaced by HIP. Estimates of harvest and hunter numbers from the Annual Questionnaire Survey for 1964-2001 are presented in Tables 6 and 7 for historical purposes. 1.4 1.6 1.8 2.0 2.2 2.4 2.6 2.8 1.4 1.6 1.8 2.0 2.2 2.4 2.6 2.8 65 68 71 74 77 80 83 86 89 92 95 98 01 EASTERN REGION CENTRAL REGION ADJUSTED AVERAGE DAILY BAG YEAR Fig. 6. Base-year adjusted indices of daily hunting success in the U.S., 1965-2002. The base year is 1969; the dashed line is the 1965-2001 average. 1.4 1.6 1.8 2.0 2.2 2.4 2.6 2.8 1.4 1.6 1.8 2.0 2.2 2.4 2.6 2.8 65 68 71 74 77 80 83 86 89 92 95 98 01 EASTERN REGION CENTRAL REGION ADJUSTED AVERAGE DAILY BAG YEAR Fig. 7. Base-year adjusted indices of seasonal hunting success in the U.S., 1965-2002. The base year is 1969; the dashed line is the 1965-2001 average. 0 20 40 60 80 100 120 140 64 69 74 79 84 89 94 99 0 100 200 300 400 500 600 700 0 20 40 60 80 100 120 140 0 100 200 300 400 500 600 700 YEAR HARVEST (THOUSANDS) HUNTERS (THOUSANDS) HUNTERS HARVEST HUNTERS HARVEST EASTERN REGION CENTRAL REGION Fig. 8. U. S. harvest of American woodcock by duck stamp purchasers, and hunter numbers, 1964-2001 (Martin 1979, and FWS unpublished data, Division of Migratory Bird Management, Laurel, Maryland). 7 Harvest Information Program Estimates of active woodcock hunters, days afield, and woodcock harvest from the 2001-02 and 2002-03 HIP surveys are provided in Table 8. In the Eastern Region woodcock hunters spent approximately 161,200 days afield and harvested 84,800 birds during 2002-03. Woodcock hunters in the Central Region spent 476,800 days afield and harvested 235,100 birds during the 2002- 03 season. Although HIP provides statewide estimates of woodcock hunter numbers (Table 8), it is not possible to develop regional estimates, due to the occurrence of some hunters visiting more than one state to hunt. ACKNOWLEDGEMENTS Rebecca Rau (FWS) mailed and processed Singing-ground Survey forms, corresponded with cooperators, keypunched portions of data, participated in the wingbee, and continued development of the new web site that allowed cooperators to submit survey data electronically. T. Nguyen and H. Bellary (FWS) played vital roles in web site and database development. Personnel from the FWS, Biological Resources Division (BRD) of the U. S. Geological Survey, Canadian Wildlife Service (CWS), and many state and provincial agencies, and other individuals assisted in collecting the Singing-ground Survey data and processing wings at the Woodcock Wingbee. Special thanks to M. Bateman (CWS), G. Haas (FWS) and S. Kelly (FWS) for help in coordinating the Singing-ground Survey. Special appreciation is extended to A. Stewart, V. Tuovila and the Michigan Department of Natural Resources for hosting the 2003 wingbee. Individuals that participated in the wingbee were: D. Dessecker and A. Bump (Ruffed Grouse Society); F. Kimmel and M. Olinde (Louisiana Department of Wildlife and Fisheries); D. McAuley (BRD); W. Palmer (Pennsylvania Game Commission); A. Stewart and V. Tuovila (Michigan DNR); R. Stonebraker (Indiana DNR); E. Oppelt and J. Bruggink (Northern Michigan U.), M. Neal (U. of Michigan-Flint) and T. Edwards, C. Horton, J. Kelley, R. Rau, P. Stinson, R. Speer, K. Sturm and L. Wolff (FWS). Thanks to the 1,198 woodcock hunters that sent in wings. M. Bateman (CWS) provided preliminary estimates of woodcock recruitment for eastern Canada. The Harvest Surveys Section of the Division of Migratory Bird Management (FWS) mailed Wing-collection Survey materials, organized wing submissions, assisted with data management, and provided Harvest Information Program estimates of woodcock harvest (special thanks to P. Padding, M. Moore, E. Martin, and J. Bezek-Balcombe). E. Martin also provided the entire dataset on woodcock harvest and hunter numbers from the Annual Questionnaire Survey. B. H. Powell (BRD) developed the computer programs for administering the Wing-collection Survey. J. Sauer (BRD) developed computer programs for calculating trends and indices from Singing-ground Survey data. W. Kendall (BRD) performed the trend analyses and assisted with interpretation. R. Rau, W. Kendall, P. Padding, J. Sauer, and M. Otto reviewed a draft of parts or all of this report and provided helpful comments. Portions of this report were copied in whole or in part from previous woodcock status reports. LITERATURE CITED Clark, E. R. 1970. Woodcock status report, 1969. U. S. Fish and Wildl. Serv., Spec. Sci. Rep.—Wildl. 133. 35pp. _____. 1972. Woodcock status report, 1971. U. S. Fish and Wildl. Serv. Spec. Sci. Rep.—Wildl. 153. 47pp. _____. 1973. Woodcock status report, 1972. U. S. Fish and Wildl. Serv. Spec. Sci. Rep.—Wildl. 169. 50pp. Coon, R. A., T. J. Dwyer, and J. W. Artmann. 1977. Identification of harvest units for the American woodcock. Proc. American Woodcock Symp. 6:147-153. Duke, G. E. 1966. Reliability of censuses of singing male woodcock. J. Wildl. Manage. 30:697-707. Dwyer, T. J., D. G. McAuley, and E. L. Derleth. 1983. Woodcock singing-ground counts and habitat changes in the northeastern United States. J. Wildl. Manage. 47:772-779. Efron, B. 1982. The jackknife, the bootstrap and other resampling plans. Society for Industrial Applied Mathematics, Philadelphia, PA. 92pp. Elden, R.C., W.V. Bevill, P.I. Padding, J.E. Frampton, and D.L. Shroufe. 2002. Pages 7-16 in J.M. Ver Steeg and R.C. Elden, compilers. Harvest Information Program: Evaluation and recommendations. Int. Assoc. of Fish and Wildl. Agencies, Migratory Shore and Upland Game Bird Working Group, Ad Hoc Committee on HIP, Washington, D. C. 100pp. Geissler, P. H. 1984. Estimation of animal population trends and annual indices from a survey of call counts or other indicators. Proceedings American Statistical Assoc., Section on Survey Research Methods, 472-477. 8 Goudy, W. H. 1960. Factors affecting woodcock spring population indexes in southern Michigan. M. S. Thesis. Michigan State Univ., E. Lansing. 44pp. Krohn, W. B., F. W. Martin, and K. P. Burnham. 1974. Band recovery distribution and survival estimates of Maine woodcock. 8pp. In Proc. Fifth American Woodcock Workshop, Athens, GA. Link, W. A., and J. R. Sauer. 1994. Estimating equations estimates of trends. Bird Populations 2:23-32. Martin, E. M. 1979. Hunting and harvest trends for migratory game birds other than waterfowl: 1964- 76. U. S. Fish and Wildl. Serv. Spec. Sci. Rep. Wildl. 218. 37pp. Martin, F. W. 1964. Woodcock age and sex determination from wings. J. Wildl. Manage. 28:287-293. _____, S. O. Williams III, J. D. Newsom, and L. L. Glasgow. 1969. Analysis of records of Louisiana-banded woodcock. Proc. 3rd Annu. Conf. Southeastern Assoc. Game and Fish Comm. 23:85- 96. Mendall, H. L., and C. M. Aldous. 1943. The ecology and management of the American woodcock. Maine Coop. Wildl. Res. Unit. Univ. Maine, Orono. 201pp. Owen, R. B., Jr., J. M. Anderson, J. W. Artmann, E. R. Clark, T. G. Dilworth, L. E. Gregg, F. W. Martin, J. D. Newsom, and S. R. Pursglove, Jr. 1977. American woodcock (Philohela minor = Scolopax minor of Edwards 1974), Pages 149-186 in G. C. Sanderson, ed. Management of migratory shore and upland game birds in North America. Int. Assoc. of Fish and Wildl. Agencies, Washington, D. C. Sauer, J. R., and J. B. Bortner. 1991. Population trends from the American Woodcock Singing-ground Survey, 1970-88. J. Wildl. Mange. 55:300-312. _____, and P. H. Geissler. 1990. Estimation of annual indices from roadside surveys. Pages 58-62 in J. R. Sauer and S. Droege, eds. Survey designs and statistical methods for the estimation of avian population trends. U. S. Fish and Wildl. Serv., Biol. Rep. 90(1). 166pp. Sepik, G. F. 1994. A woodcock in the hand. Ruffed Grouse Society, Coraopolis, PA. 12pp. Straw, J. A., D. G. Krementz, M. W. Olinde, and G. F. Sepik. 1994. American woodcock. Pages 97-114 in T. C. Tacha and C. E. Braun, eds. Migratory Shore and Upland Game Bird Management in North America. Int. Assoc. of Fish and Wildl. Agencies, Washington, D. C. U. S. Fish and Wildlife Service. 1990. American woodcock management plan. U. S. Fish and Wildl. Serv., Washington, D. C. 11pp. Whitcomb, D. A. 1974. Characteristics of an insular woodcock population. Mich. Dept. Nat Resour., Wildl. Div. Rep. 2720. 78pp. 9 Table 1. Trends (% change per yeara and 90% confidence interval) in the number of American woodcock heard in the Singing-ground Survey as determined by the estimating equations technique (Link and Sauer 1994), 1968-2003. State, 2002-2003 1993-2003 1968-2003 Province No. of nc % change 90% CI n % change 90% CI n % change 90% CI CT 4 2 0.0 0.0 0.0 4 17.7 0.0 35.5 9 -10.0 ** d -16.5 -3.5 DE 2 0 2 -10.0 * -19.9 -0.1 2 4.2 -11.3 19.7 ME 44 29 23.1 * 1.0 45.1 53 -0.6 -2.0 0.8 64 -2.3 *** -3.1 -1.4 MD 6 2 -26.7 -110.8 57.4 6 -15.3 -43.0 12.5 21 -10.8 *** -17.2 -4.4 MA 7 6 -7.1 -43.8 29.6 9 5.2 ** 1.0 9.5 20 -4.1 ** -7.4 -0.7 NB 32 25 -2.1 -17.3 13.2 51 0.9 -1.5 3.3 62 -0.7 -1.9 0.5 NH 13 12 34.0 -8.6 76.7 13 0.7 -3.5 4.9 18 0.5 -2.2 3.3 NJ 7 2 28.7 -1.8 59.2 5 -4.1 -13.6 5.5 17 -10.1 *** -13.5 -6.8 NY 59 38 12.0 -14.7 38.6 70 -2.9 ** -4.9 -0.9 105 -2.8 *** -3.8 -1.8 NS 30 19 1.9 -23.9 27.7 38 1.1 -1.9 4.0 56 -0.3 -1.6 0.9 PA 34 13 -12.8 -41.9 16.3 26 -1.6 -7.2 4.1 56 -4.7 *** -6.9 -2.5 PEI 0 0 7 -2.8f -7.2 1.6 12 -1.4g -2.9 0.1 QUE 26 0 13 -2.9 ** -5.1 -0.7 54 -1.8 -4.9 1.2 RI 1 0 2 -15.7 *** -23.1 -8.2 VT 17 10 29.6 -11.2 70.3 18 0.7 -2.3 3.6 21 -1.5 -3.2 0.1 VA 26 4 17.0 -145.5 179.4 12 -10.7 * -20.2 -1.2 47 -10.7 *** -13.9 -7.4 WV 27 8 27.7 -58.1 113.6 14 -5.7 ** -9.8 -1.6 43 -2.7 *** -4.3 -1.0 Eastern 335 173 6.4 -4.9 17.7 341 -1.3 ** -2.4 -0.3 609 -2.3 *** -2.8 -1.8 IL 14 0 4 13.0 -17.2 43.2 23 23.4 -10.2 57.1 IN 17 2 0.0 0.0 0.0 7 -12.4 -28.3 3.6 38 -6.1 * -11.5 -0.7 MBe 18 9 31.7 -3.8 67.3 20 -4.0 * -7.8 -0.2 20 -3.5 ** -6.2 -0.9 MI 89 43 11.3 -6.1 28.6 109 -1.2 -2.6 0.1 143 -1.7 *** -2.5 -0.8 MN 72 46 5.5 -13.3 24.3 79 -0.4 -2.2 1.4 99 -1.1 ** -2.0 -0.3 OH 30 9 -9.9 -62.5 42.7 27 -10.6 ** -18.7 -2.6 55 -6.3 *** -10.0 -2.7 ON 34 0 87 -3.2 -7.9 1.5 136 -1.6 *** -2.3 -0.9 WI 69 41 7.9 -12.1 27.8 71 -1.7 -3.7 0.3 100 -1.9 *** -2.7 -1.0 Central 343 151 8.7 -1.7 19.2 404 -1.6 *** -2.5 -0.7 614 -1.8 *** -2.2 -1.3 Continent 635 324 7.7 * 0.7 14.8 745 -1.5 *** -2.2 -0.8 1223 -1.9 *** -2.3 -1.6 a Mean of weighted route trends within each state, province or region. To estimate the total percent change over several years, use: (100((% change/100)+1)y)-100 where y is the number of years. Note: extrapolating the estimated trend statistic (% change per year) over time (e.g., 30 years) may exaggerate the total change over the period. b Total number of routes surveyed in 2003 for which data were received by 30 May. c Number of comparable routes (2002 versus 2003) with at least 2 non-zero counts. d Indicates slope is significantly different from zero: * P<0.10, ** P<0.05. *** P <0.01; significance levels are approximate for states where n<10. e Manitoba began participating in the Singing-ground Survey in 1990. f Data were not received from PEI for the 2002 and 2003 surveys. Trend estimate is for the period 1993-2001. g Data were not received from PEI for the 2002 and 2003 surveys. Trend estimate is for the period 1968-2001. 10 1985 1.38 0.60 3.81 1.33 1.99 3.98 2.65 1.95 3.59 2.16 1.48 2.88 3.52 0.64 2.10 0.87 0.89 2.46 0.39 0.67 -- 4.89 3.66 1.47 5.02 3.03 2.98 2.53 1984 1.61 0.59 3.75 1.32 2.54 3.67 2.50 2.73 2.84 2.15 1.90 3.95 2.88 1.92 2.66 1.72 0.94 2.56 0.22 0.82 -- 4.68 3.06 1.70 4.88 3.28 2.78 2.59 1983 2.44 1.63 3.75 1.68 1.44 4.54 2.81 2.31 3.53 2.25 1.80 3.46 3.67 2.18 2.62 1.20 1.14 2.70 0.19 0.84 -- 4.25 3.44 1.83 4.64 3.00 2.87 2.65 1982 3.18 -- 2.92 2.52 1.93 4.33 2.40 1.99 3.06 1.80 1.59 2.18 2.96 3.19 1.79 1.55 1.10 2.44 0.13 0.80 -- 4.87 3.78 1.46 4.49 2.99 2.66 2.71 1981 2.57 -- 4.24 2.56 2.26 4.16 4.09 1.96 3.78 2.02 1.96 2.05 3.08 0.78 2.38 1.65 1.25 2.74 0.15 1.09 -- 4.61 4.19 2.03 5.92 3.05 3.17 2.78 1980 1.89 -- 3.87 3.00 2.20 4.12 3.91 2.55 4.16 2.18 1.98 2.72 3.96 1.35 2.67 1.68 0.91 2.77 0.10 1.05 -- 5.51 4.57 1.78 6.38 3.58 3.23 2.84 1979 2.16 0.41 4.40 2.43 3.06 4.61 3.22 4.12 3.61 2.29 2.17 3.67 3.62 1.35 3.00 1.98 1.10 2.91 0.08 1.43 -- 5.60 4.11 1.80 6.24 4.19 3.43 2.91 1978 2.18 0.48 4.03 3.08 2.80 4.15 3.14 2.41 3.20 2.82 1.91 2.96 3.57 0.78 3.11 1.81 0.76 2.63 0.07 1.15 -- 5.69 4.17 2.33 6.49 4.29 3.48 2.98 1977 3.60 0.51 4.37 2.90 2.40 5.80 3.12 4.16 4.04 2.44 2.40 3.70 2.92 -- 4.07 2.38 1.09 3.02 0.07 1.28 -- 5.38 4.16 2.93 6.00 4.07 3.50 3.05 1976 3.08 0.38 4.84 3.06 3.09 4.66 3.79 3.77 3.93 2.42 2.42 4.19 2.64 2.35 3.40 2.49 1.08 2.90 0.05 1.32 -- 5.88 4.20 2.57 5.51 3.75 3.42 3.12 1975 5.63 1.20 5.46 4.51 2.33 6.35 3.06 6.23 3.98 2.74 2.50 4.99 3.82 2.35 3.75 3.00 1.24 3.37 0.06 1.34 -- 6.39 4.17 2.37 5.74 3.91 3.60 3.19 1974 5.31 0.72 5.11 4.26 3.99 5.60 3.61 8.36 4.76 3.20 2.23 3.27 3.82 3.03 3.22 3.53 1.08 3.42 0.03 1.39 -- 6.35 4.80 3.16 6.53 4.03 3.66 3.27 1973 5.48 0.77 5.15 6.04 4.94 5.05 2.63 8.41 4.52 2.56 3.09 2.46 3.22 4.06 3.28 2.41 1.12 3.24 0.03 1.89 -- 5.47 4.13 2.46 6.09 3.94 3.52 3.34 1972 7.49 0.46 4.81 5.45 3.67 5.69 3.38 5.98 4.49 2.64 2.86 3.07 4.26 4.06 3.71 3.35 1.40 3.49 0.03 1.85 -- 5.33 3.63 2.97 6.82 3.87 3.60 3.42 1971 6.14 0.40 5.11 6.51 4.99 5.46 2.68 10.09 4.79 2.76 3.22 5.26 4.51 5.32 3.29 3.85 1.15 3.63 0.02 1.53 -- 5.63 4.27 3.55 6.15 4.05 3.66 3.50 1970 7.92 0.55 5.63 7.00 4.26 5.57 3.31 8.13 4.25 2.23 3.71 2.84 -- 2.82 4.30 4.83 1.19 3.72 0.02 1.98 -- 5.83 3.98 3.50 6.47 4.57 3.80 3.58 1969 7.94 0.48 5.36 7.89 3.63 5.25 2.87 6.50 5.54 2.62 3.38 3.81 -- 2.83 2.53 4.66 1.66 3.80 -- 2.08 -- 6.12 4.64 -- 6.81 4.22 3.90 3.67 Year 1968 --b 0.62 5.19 8.94 -- -- -- 7.52 5.03 3.61 3.67 -- -- -- -- -- 1.48 3.92 -- 2.40 -- 6.26 -- -- 6.22 4.28 3.91 3.76 Table 2. Breeding population indices for American woodcock from the Sing-ground Survey, 1968-2003. These indices are based on the 1968-2003 trend and should be used for exploratory data analysis only; observed patterns should be verified using trend estimation methods (Sauer and Geissler 1990). State, Province or Region Eastern Region CTa DEa ME MD MA NB NH NJ NY NS PA PEIa QUEa RIa VT VA WV Region Central Region IL IN MB MI MN OH ON WI Region Continent a Annual indices are unreliable due to small sample size. b Insufficient data. 11 2003 0.30 0.72 2.67 0.19 1.46 4.75 3.97 0.33 1.98 2.16 0.77 -- 2.54 0.02 2.08 0.16 0.62 1.78 6.96 0.29 1.93 3.53 2.95 0.47 4.18 2.32 2.16 1.66 2002 0.31 0.72 2.46 0.29 1.32 3.91 3.63 0.38 1.80 2.06 0.80 0.87 2.58 0.05 1.83 0.18 0.51 1.66 4.46 0.26 1.45 3.42 2.72 0.49 5.94 2.19 2.04 1.70 2001 0.33 0.67 2.58 0.63 1.28 4.89 3.41 0.58 2.02 2.56 0.76 3.05 2.29 -- 2.22 0.19 0.61 1.81 6.30 0.46 2.36 3.35 3.55 0.54 3.88 2.34 2.37 1.74 2000 0.89 0.93 2.99 0.35 1.38 4.54 3.22 0.59 1.97 2.75 0.61 2.84 2.43 -- 3.47 0.23 0.74 1.84 3.81 0.40 1.88 3.61 3.45 0.63 4.66 2.55 2.29 1.78 1999 1.32 0.42 3.06 0.34 2.05 4.96 4.69 0.84 2.19 2.32 0.92 2.42 2.95 -- 2.57 0.26 0.65 2.02 2.75 0.47 1.73 3.46 3.18 0.56 3.96 2.78 2.31 1.83 1998 0.62 1.42 2.41 0.24 1.35 3.99 3.81 0.67 2.24 2.32 1.17 2.86 2.35 -- 2.56 0.25 0.62 1.86 -- 0.73 1.80 4.30 3.22 0.72 3.99 2.30 2.44 1.87 1997 0.65 0.72 2.56 0.51 1.47 4.80 4.14 0.20 2.17 1.99 1.06 2.66 2.29 0.08 2.29 0.34 0.72 1.89 1.69 0.35 1.47 3.61 2.60 0.61 4.00 2.36 1.87 1.91 1996 0.74 0.72 2.31 0.44 1.37 3.93 3.76 0.97 2.19 2.61 0.99 3.06 1.19 -- 1.75 0.25 0.65 1.66 3.78 0.45 2.49 3.70 2.97 0.84 3.47 2.52 2.33 1.96 1995 0.81 -- 3.02 0.31 1.05 4.34 4.76 0.81 2.35 2.54 1.21 2.69 3.28 -- 2.32 0.29 1.04 2.17 1.21 0.53 2.77 3.86 3.29 0.80 4.75 2.40 2.43 2.00 1994 0.64 -- 2.84 0.52 1.44 5.14 2.42 0.34 2.26 2.07 0.65 2.31 2.75 -- 2.06 0.38 0.60 1.80 1.33 0.51 2.47 3.58 3.03 0.79 3.80 2.38 2.31 2.05 1993 0.51 -- 3.22 0.58 1.28 5.34 2.85 0.80 2.26 2.74 1.31 2.27 3.54 -- 2.04 0.49 0.69 2.16 1.27 0.59 3.43 3.95 3.47 0.95 4.38 2.55 2.63 2.10 1992 0.62 0.24 2.98 0.30 1.51 3.97 2.29 0.85 2.79 2.50 1.23 2.40 2.98 -- 1.93 0.43 0.78 2.07 1.06 0.55 2.61 3.93 3.26 0.92 4.85 2.59 2.51 2.15 1991 0.90 0.39 3.58 0.72 1.83 4.19 3.94 1.03 3.27 2.28 1.68 2.45 3.54 0.20 2.95 0.55 0.78 2.43 0.79 0.71 -- 5.50 3.84 1.04 5.05 3.25 2.98 2.20 1990 0.85 0.72 2.85 0.87 1.52 4.38 2.87 1.08 3.03 1.85 1.52 3.29 2.89 -- 3.01 0.57 0.85 2.21 0.52 0.69 -- 4.68 4.11 1.32 5.08 3.21 2.84 2.25 1989 0.98 -- 4.15 1.10 1.67 5.53 3.31 1.59 2.53 2.69 1.15 4.02 3.73 0.96 3.15 0.58 0.80 2.37 0.60 0.62 -- 4.79 3.57 1.01 5.39 3.29 2.82 2.30 1988 2.33 -- 4.06 0.97 2.12 4.26 3.19 1.67 3.27 2.47 1.60 4.23 2.51 0.96 3.39 0.66 0.79 2.39 0.49 0.61 -- 5.02 4.13 1.48 5.10 3.55 2.98 2.36 1987 0.92 -- 4.31 0.92 2.12 3.97 3.23 2.22 2.81 2.27 1.61 2.64 3.53 -- 2.86 0.93 0.99 2.53 0.48 0.65 -- 4.59 3.69 1.24 5.17 3.55 2.98 2.41 Year 1986 2.00 --b 3.94 1.17 2.04 3.33 4.51 1.90 3.06 2.53 1.70 3.78 3.39 0.64 2.66 0.90 0.86 2.50 0.32 0.90 -- 4.96 3.86 1.14 4.94 3.56 2.97 2.47 Table 2. Continued. State, Province or Region Eastern Region CTa DEa ME MD MA NB NH NJ NY NS PA PEIa QUEa RIa VT VA WV Region Central Region IL IN MB MI MN OH ON WI Region Continent a Annual indices are unreliable due to small sample size. b Insufficient data. 12 Table 3. Distribution of U.S. hunters contacted and hunters that submitted woodcock wings in the 2002-03 Wing-collection Survey. State of No. of hunters No. of hunters that Percent that residence contacted submitted wings submitted wings AL 32 0 0 AR 29 0 0 CT 207 21 10 DE 24 1 4 FL 98 0 0 GA 77 6 8 IL 161 4 2 IN 138 29 21 IA 73 6 8 KS 16 0 0 KY 32 2 6 LA 194 20 10 ME 403 119 30 MD 84 10 12 MA 421 70 17 MI 777 242 31 MN 535 75 14 MS 23 0 0 MO 145 13 9 NE 39 0 0 NH 224 77 34 NJ 180 37 21 NY 459 94 20 NC 104 5 5 ND 7 0 0 OH 190 20 11 OK 37 0 0 PA 495 61 12 RI 49 5 10 SC 78 10 13 TN 72 3 4 TX 71 3 4 VT 179 68 38 VA 167 18 11 WV 42 16 38 WI 941 163 17 Total 6,803 1,198 18 13 Table 4. Numbers of woodcock wings received from hunters, and indices of recruitment in the U.S. Recruitment indices for individual states were calculated as the ratio of immatures per adult female. The regional indices for 2002 were calculated as the average of the state values, adjusted for comparability with the 1963-2001 average. Recruitment indices were not calculated for states where the sample of wings was <125. State or Wings received Region of Total Adult females Immatures Recruitment index harvest 1963-01 2002 1963-01 2002 1963-01 2002 1963-01 2002 Eastern Region CT 13,149 89 2,909 19 8,066 57 2.8 DE 418 3 54 0 294 2 5.4 FL 660 0 150 0 410 0 2.7 GA 2,927 29 901 10 1,270 7 1.4 ME 73,296 969 21,567 301 36,667 486 1.7 1.6 MD 3,843 42 958 16 2,150 19 2.2 MA 19,253 428 5,820 142 9,550 182 1.6 1.3 NH 27,456 702 8,876 227 12,697 333 1.4 1.5 NJ 24,357 222 5,660 60 14,331 105 2.5 1.8 NY 51,328 748 16,975 251 23,890 317 1.4 1.3 NC 3,007 65 890 30 1,489 30 1.7 PA 27,589 351 8,705 109 12,777 146 1.5 1.3 RI 2,246 13 420 2 1,519 8 3.6 SC 2,312 105 712 25 1,102 44 1.5 VT 20,311 457 6,513 173 9,494 197 1.5 1.1 VA 3,948 121 942 53 2,274 44 2.4 WV 5,156 76 1,572 22 2,611 32 1.7 Region 281,256 4,420 83,624 1,440 140,591 1,872 1.7 1.4 Central Region AL 910 0 243 0 425 0 1.7 AR 515 0 165 0 207 0 1.3 IL 1,289 18 293 4 727 13 2.5 IN 6,819 138 1,717 37 3,802 61 2.2 1.6 IA 922 36 311 12 407 16 1.3 KS 44 0 9 0 22 0 a KY 1,010 23 238 6 524 15 2.2 LA 28,894 318 6,479 64 18,711 211 2.9 3.3 MI 100,620 2,239 32,536 677 50,320 1122 1.5 1.7 MN 28,604 538 9,708 224 12,840 183 1.3 0.8 MS 1,719 0 488 0 875 0 1.8 MO 2,703 84 668 22 1,360 45 2.0 NE 13 0 5 0 6 0 a OH 13,648 118 4,137 34 6,476 59 1.6 1.8 OK 170 2 38 0 89 2 2.3 TN 1,008 10 250 2 517 4 2.1 TX 986 1 261 1 501 0 1.9 WI 62,678 1,057 20,444 353 30,429 492 1.5 1.4 Region 252,552 4,582 77,990 1,436 128,238 2,478 1.6 1.6 14 Table 5. State and regional indices of daily and seasonal woodcock hunting success in the U.S. during 2001 and 2002. State and regional indices were calculated for states represented by >10 hunters that participated in the Wing-collection Survey both years. Regional indices were weighted as described by Clark (1970). State of No. of successful No. of successful hunts Woodcock bagged Woodcock per successful hunt Woodcock per season harvest hunters 2001 2002 2001 2002 2001 2002 2001 2002 Eastern Region CT 8 26 21 49 45 DE 1 2 1 4 3 GA 4 5 11 6 26 ME 84 391 363 875 795 2.2 2.2 10.4 9.5 MD 6 12 6 25 12 MA 33 132 152 247 275 1.9 1.8 7.5 8.3 NH 46 266 249 576 512 2.2 2.1 12.5 11.1 NJ 16 63 60 132 135 2.1 2.3 8.3 8.4 NY 60 278 290 507 559 1.8 1.9 8.5 9.3 NC 4 30 28 68 61 PA 39 143 129 319 272 2.2 2.1 8.2 7.0 RI 4 7 7 10 12 SC 9 27 43 60 99 VT 40 170 171 309 331 1.8 1.9 7.7 8.3 VA 7 45 45 104 86 2.2 2.2 12.6 10.1 WV 6 14 13 22 26 Region 367 1,611 1,589 3,313 3,249 1.8 1.9 6.9 6.6 Central Region IA 6 21 20 33 35 IL 3 6 9 10 15 IN 14 47 50 94 93 2.0 1.9 6.7 6.6 KY 2 15 12 31 23 LA 13 83 107 220 306 2.7 2.9 16.9 23.5 MI 186 1,076 955 2,224 1,932 2.1 2.0 12.0 10.4 MN 64 311 239 634 490 2.0 2.1 9.9 7.7 MO 5 22 27 50 60 OH 12 78 51 168 112 2.2 2.2 14.0 9.3 TN 2 14 5 25 7 TX 1 4 1 6 1 WI 120 545 406 1,111 840 2.0 2.1 9.3 7.0 Region 428 2,222 1,882 4,606 3,914 2.1 2.1 10.0 11.0 15 1982 10,981 1,637 419 4,355 4,437 18,376 7,788 12,951 6,856 15,450 33,513 8,959 33,329 1,283 4,459 6,042 2,770 2,874 176,479 2,874 4,802 12,707 7,131 4,015 952 3,864 112,186 60,222 39,761 8,926 4,076 862 246 12,273 904 98 4,739 5,893 43,688 330,219 1981 12,245 1,714 193 4,155 5,440 25,390 10,743 21,688 12,099 13,901 56,536 13,228 39,689 1,261 7,298 7,777 5,854 1,494 240,705 2,425 5,445 13,108 7,006 3,339 831 3,892 211,441 56,733 32,205 9,777 6,291 1,559 90 13,216 1,316 58 4,448 5,172 64,617 442,969 1980 15,217 2,929 184 5,251 6,691 30,132 8,329 20,109 12,780 27,108 46,196 6,973 23,740 1,186 7,503 6,185 4,691 3,230 228,433 5,132 6,389 11,346 10,965 6,904 785 2,883 327,751 71,999 52,627 21,742 8,516 1,055 326 14,931 1,265 161 2,822 16,900 77,072 641,571 1979 13,488 3,210 343 3,956 5,977 33,452 5,267 20,114 15,268 23,559 71,113 7,252 35,807 2,708 6,199 6,234 7,696 2,563 264,205 4,788 7,647 13,800 9,309 5,460 510 3,695 146,576 72,738 48,432 11,904 6,540 378 0 12,153 908 55 6,282 10,392 75,913 437,480 1978 15,905 4,349 399 5,667 6,747 36,279 11,261 23,949 19,860 30,933 48,286 9,897 37,391 2,213 8,926 8,567 6,628 3,418 280,675 4,514 6,081 19,533 8,396 8,662 935 4,963 214,793 98,260 53,865 11,042 12,929 1,203 292 12,112 1,384 0 9,375 6,689 96,926 571,954 1977 17,792 2,535 1,084 6,596 9,911 32,347 7,839 21,185 12,936 24,700 66,796 9,186 33,409 1,216 7,716 6,367 5,043 4,706 271,364 7,975 20,586 14,347 6,384 5,587 2,419 1,874 130,271 122,600 44,676 10,540 9,438 536 0 14,575 2,894 46 3,668 15,916 114,422 528,754 1976 20,134 4,165 191 8,881 8,623 35,440 9,883 28,492 16,399 37,937 59,342 8,936 43,550 2,624 7,288 16,443 9,421 3,868 321,617 5,796 8,162 10,481 10,583 5,025 1,115 4,260 121,740 128,568 18,019 17,651 8,733 540 150 14,891 2,779 135 4,082 12,207 125,453 500,370 1975 16,181 4,593 450 4,909 6,714 40,846 7,568 33,971 16,109 31,377 72,259 8,899 37,573 2,078 8,010 11,117 10,162 2,273 315,089 6,909 9,475 11,302 15,372 2,619 892 3,126 59,758 157,417 35,806 10,925 6,545 394 352 13,955 4,648 135 4,648 10,471 105,416 460,165 1974 13,547 3,772 500 5,402 10,999 48,123 8,530 31,648 16,754 42,308 83,600 14,617 45,170 3,457 9,363 9,234 5,392 2,345 354,761 3,532 6,811 11,480 12,163 4,584 2,297 2,330 67,492 114,106 25,290 11,082 6,145 520 378 16,906 2,117 0 5,724 5,516 101,895 400,368 1973 18,729 4,759 135 6,012 5,997 34,765 8,475 37,623 13,752 42,975 82,810 8,186 43,090 2,472 5,660 8,723 5,911 1,892 331,966 3,035 8,451 8,531 6,993 3,416 2,019 1,727 47,436 119,723 20,682 12,775 6,376 477 325 13,274 2,233 124 3,544 6,817 58,405 326,363 1972 12,953 2,057 975 5,439 8,186 35,762 4,382 34,958 12,098 23,906 77,935 9,970 49,599 2,238 9,280 7,061 4,461 4,305 305,565 6,721 8,139 10,509 8,797 2,945 1,131 790 134,592 95,512 26,589 12,997 2,880 489 825 10,511 1,696 82 4,257 5,428 82,265 417,155 1971 14,619 4,109 1,232 4,192 6,208 33,923 8,334 25,363 12,556 39,662 80,334 9,254 57,311 4,049 7,783 10,384 4,470 1,715 325,498 3,985 4,755 9,684 12,159 756 735 1,389 73,272 106,387 20,745 8,409 3,482 299 395 18,031 1,610 0 2,926 5,678 65,915 340,612 1970 23,430 3,626 360 6,868 5,579 33,171 9,781 32,197 9,520 28,030 74,873 9,181 54,754 2,585 9,050 11,459 4,120 1,454 320,038 5,424 3,723 7,589 6,293 1,085 1,073 788 95,777 77,671 20,721 9,534 4,294 1,333 74 13,136 836 93 1,727 4,353 90,768 346,292 1969 18,815 3,043 629 4,568 3,645 25,805 10,082 32,221 13,099 28,207 83,081 6,995 42,136 3,092 8,716 5,173 6,967 467 296,741 7,432 2,619 9,705 6,210 861 885 2,382 105,955 54,402 14,182 8,820 3,919 192 0 19,445 1,977 84 2,626 6,234 44,894 292,824 1968 15,772 3,798 330 5,799 6,226 38,617 9,127 29,330 15,483 24,622 76,528 9,612 30,917 2,086 8,741 4,553 4,342 1,213 287,076 4,114 3,193 2,083 3,884 483 629 692 75,271 42,244 7,948 6,337 1,529 33 348 15,345 535 32 2,602 2,804 27,794 197,900 1967 12,238 1,767 553 3,999 4,672 27,726 7,011 20,532 8,212 20,791 75,869 8,308 39,811 1,758 4,622 9,133 3,988 926 251,916 5,182 6,354 4,109 5,048 1,668 278 702 62,869 73,928 16,591 6,982 3,245 515 0 10,190 2,866 341 1,564 2,329 40,282 245,043 1966 13,137 2,210 747 3,453 2,594 25,350 3,103 25,056 9,490 14,094 49,459 9,042 18,236 1,584 3,000 7,178 3,260 413 191,406 5,458 2,684 4,133 4,145 222 268 667 53,842 68,661 9,952 4,556 2,392 56 441 9,776 386 90 1,097 4,972 36,190 209,988 1965 16,261 1,879 240 3,501 2,341 24,158 3,318 28,156 11,259 13,437 44,261 5,219 29,119 1,453 2,707 3,421 2,317 584 193,361 1,626 3,683 3,124 2,921 433 271 427 31,855 62,910 4,929 3,095 2,634 104 152 13,011 57 0 856 1,968 48,905 182,961 1964 11,928 1,804 91 2,126 2,893 26,115 3,025 19,696 6,488 11,108 38,625 6,790 19,340 541 8,552 4,233 2,648 186 166,189 2,224 3,952 4,015 4,761 1,563 547 737 71,744 90,662 10,795 7,390 2,646 253 44 16,905 327 100 488 8,115 58,934 286,202 Table 6. State and regional estimates of the number of American woodcock bagged (retrieved kill) by duck stamp buyers in the U.S., 1964-2001. Data from Martin (1979) and unpublished FWS administrative reports. Information was not available to reassign hunters from the District of Columbia to state of harvest. State or Region CT DE DC FL GA ME MD MA NH NJ NY NC PA RI SC VT VA WV Eastern Region AL AR IL IN IA KS KY LA MI MN MS MO NE ND OH OK SD TN TX WI Central Region 16 2001 3,748 392 121 4,924 1,990 14,087 1,973 5,511 3,792 4,623 5,593 4,226 11,866 187 4,626 3,446 3,280 923 75,311 124 231 3,256 4,974 948 442 3,008 7,026 35,290 16,044 955 3,646 823 238 6,280 1,318 117 1,341 2,727 14,253 103,041 2000 1,996 275 164 1,007 2,211 18,680 2,041 0 3,717 2,478 5,272 1,868 11,008 404 3,455 4,549 3,605 130 62,859 1,620 663 3,486 1,440 1,718 838 449 32,409 48,231 22,638 1,624 2,392 2,247 0 5,239 811 257 1,529 764 19,459 147,814 1999 5,306 0 214 787 1,503 15,028 2,099 5,818 2,668 4,286 8,886 4,403 8,020 619 5,061 3,711 10,025 2,221 80,654 80 4,053 3,071 2,520 895 3,303 979 23,066 61,992 23,456 326 1,684 0 0 6,273 1,264 0 4,293 2,576 22,709 162,540 1998 2,180 595 145 329 2,491 19,647 1,849 4,200 3,793 8,841 12,421 3,093 9,183 719 3,521 787 2,522 158 76,475 1,815 2,004 4,594 5,831 2,909 317 2,044 12,904 67,411 24,988 886 3,718 771 432 7,953 1,738 340 1,839 6,038 30,443 178,975 1997 2,718 477 216 975 898 15,479 2,034 5,233 3,037 4,131 10,735 2,962 9,550 203 2,521 2,688 3,234 0 67,091 1,117 1,299 4,152 3,022 2,478 792 412 12,645 45,417 29,865 5,627 2,483 2,185 0 6,487 585 21 9,116 6,002 30,714 164,419 1996 5,496 508 406 535 3,764 12,322 1,797 7,309 5,235 2,081 11,877 3,305 11,087 965 6,369 8,165 18,262 63 99,546 796 2,304 2,736 4,708 1,236 597 3,081 24,931 54,086 42,001 1,265 2,161 712 842 6,675 355 63 1,648 2,715 30,819 183,731 1995 6,209 641 137 187 1,989 18,227 3,253 12,216 5,023 4,338 11,955 2,803 7,918 538 5,244 4,274 2,943 389 88,282 599 274 5,023 4,978 1,864 883 1,708 67,428 70,329 31,409 2,708 2,043 162 255 4,721 630 41 2,813 2,985 28,706 229,559 1994 7,260 739 114 6,363 2,348 21,599 2,715 7,773 4,437 5,827 12,701 4,669 10,784 1,463 3,863 3,755 2,317 522 99,254 150 1,621 6,846 4,064 1,306 121 1,262 40,285 62,462 27,183 1,212 993 656 85 6,324 251 0 2,497 5,757 36,820 199,895 1993 4,630 317 149 1,422 2,843 14,551 3,198 8,378 4,158 7,882 10,199 2,789 13,476 481 3,032 2,384 1,898 35 81,822 356 248 5,849 2,914 1,172 225 1,265 44,096 54,058 20,260 889 1,962 153 569 7,950 524 0 1,009 2,109 32,076 177,684 1992 4,163 2,322 303 307 971 23,779 4,095 13,279 5,915 2,301 16,287 4,084 13,705 888 1,919 3,099 5,987 125 103,530 1,228 10,166 11,471 3,011 2,251 61 2,678 29,494 60,230 27,929 1,531 3,578 828 55 9,786 2,898 84 1,146 1,775 38,255 208,455 1991 5,174 1,281 120 741 2,513 20,550 3,506 13,751 6,616 7,821 12,371 4,750 11,719 2,061 1,759 4,411 4,343 650 104,259 651 1,780 9,804 2,431 1,972 612 1,932 37,289 66,833 35,917 994 3,349 303 701 8,518 716 1,521 1,763 3,775 36,968 217,829 1990 6,890 1,358 128 1,906 2,758 13,489 1,910 11,109 5,014 7,542 15,384 3,880 18,089 1,270 2,287 6,004 5,148 379 104,544 812 1,470 8,031 2,927 2,467 421 2,523 39,735 61,878 36,136 3,401 3,550 1,072 150 8,464 1,078 0 4,865 3,940 44,019 226,939 1989 5,884 3,230 0 2,405 2,863 16,093 6,574 13,083 5,808 10,900 20,041 5,159 15,220 491 4,518 5,098 4,684 854 122,904 1,886 1,220 8,935 6,685 5,457 616 3,343 66,615 50,856 45,396 2,065 6,279 1,124 156 15,165 2,268 219 3,996 5,434 42,965 270,680 1988 7,059 912 365 1,106 3,657 13,760 5,821 10,871 7,201 9,704 18,414 3,502 11,569 1,455 3,487 9,539 6,491 751 115,664 1,984 654 3,031 4,428 1,269 174 985 43,689 52,266 45,579 1,927 3,453 701 580 4,244 1,046 0 1,596 8,216 46,452 222,274 1987 7,374 1,694 370 4,270 2,203 27,404 9,887 15,793 6,567 10,885 27,073 5,373 21,354 2,530 6,569 7,198 6,256 409 163,209 3,298 5,682 7,672 8,194 3,897 2,108 1,287 123,958 70,062 71,966 2,891 7,818 481 0 5,866 1,097 580 3,506 9,498 61,375 391,236 1986 4,838 901 45 2,195 3,851 22,652 3,647 14,752 7,800 7,175 20,208 7,058 20,326 639 5,095 5,883 3,002 1,664 131,730 1,942 3,188 6,898 5,882 3,656 1,681 2,848 110,850 68,752 34,505 3,370 5,917 1,125 0 8,888 1,321 769 3,269 11,479 59,843 336,183 1985 6,319 849 65 2,608 2,988 17,314 7,221 17,139 7,420 8,804 17,945 7,299 16,214 1,988 2,094 4,439 7,112 578 128,394 2,108 6,636 7,285 4,921 3,888 1,260 1,991 78,532 56,885 28,678 6,388 4,087 364 754 8,719 2,043 0 3,235 7,922 42,360 268,056 1984 8,989 2,771 250 3,414 6,927 21,023 11,800 12,131 7,912 15,962 27,605 5,725 23,927 843 5,211 6,172 6,891 1,650 169,204 2,849 2,473 7,266 5,566 2,447 916 3,983 106,912 64,823 26,304 6,423 6,329 1,035 474 7,381 1,346 0 2,115 9,209 64,972 322,823 1983 9,378 1,210 0 5,661 5,231 19,155 6,812 11,937 4,701 11,391 27,789 5,859 27,340 1,915 7,137 4,651 2,783 3,293 156,244 2,408 2,584 8,047 4,865 2,731 1,164 3,412 196,201 55,615 35,346 4,839 4,133 3,257 264 12,941 1,408 275 2,479 8,510 44,112 394,591 Table 6. continued State or Region CT DE DC FL GA ME MD MA NH NJ NY NC PA RI SC VT VA WV Eastern Region AL AR IL IN IA KS KY LA MI MN MS MO NE ND OH OK SD TN TX WI Central Region 17 1982 3,749 778 76 828 1,509 5,058 2,037 6,169 2,967 3,993 13,536 2,676 14,972 578 1,770 2,152 1,434 564 64,846 798 1,720 3,711 2,102 1,463 512 872 19,233 14,406 11,791 1,424 1,839 435 227 4,490 338 87 1,239 1,824 16,851 85,362 1981 5,574 903 96 1,295 1,731 5,881 3,259 6,544 3,823 4,997 16,894 3,011 13,374 516 2,021 2,322 1,507 464 74,571 761 1,598 3,883 2,165 1,742 184 932 21,651 14,650 11,164 2,100 2,509 387 57 4,513 392 81 1,461 1,591 18,544 90,365 1980 5,173 905 73 1,366 2,091 5,920 2,117 5,609 3,526 4,983 15,150 2,470 13,522 397 2,569 2,197 1,447 513 70,030 1,067 1,976 4,340 2,485 2,443 419 706 27,508 17,000 17,098 2,544 2,780 325 137 4,770 444 73 1,176 3,068 22,111 112,470 1979 4,836 913 100 1,104 1,927 5,745 2,752 6,409 4,244 6,209 18,405 1,716 16,136 772 1,633 2,060 2,017 551 77,527 1,244 2,356 4,437 2,682 2,089 469 1,182 19,529 19,878 14,355 2,168 2,200 278 28 4,469 504 47 2,252 2,612 21,245 104,024 1978 5,614 1,439 199 1,382 1,746 6,910 3,115 6,026 4,448 8,227 17,330 2,815 19,591 763 2,603 2,460 1,986 501 87,155 1,406 1,642 6,729 2,794 4,110 510 1,750 24,041 20,271 16,925 2,683 3,288 564 182 3,678 776 31 2,378 2,556 20,489 116,803 1977 6,209 1,370 321 1,413 1,933 6,205 2,584 7,086 4,379 7,188 19,628 2,373 16,814 629 2,201 1,791 1,609 492 84,225 1,703 3,035 3,912 2,152 3,011 1,470 904 17,795 23,151 14,155 2,579 3,041 357 0 4,971 856 46 1,313 3,730 27,274 115,455 1976 5,987 1,675 93 2,042 1,428 6,397 3,675 7,824 4,761 8,337 17,345 2,857 18,700 990 1,987 2,782 2,132 457 89,469 1,459 2,666 4,108 2,717 2,619 796 1,249 16,463 30,685 7,857 2,864 2,387 270 150 5,623 898 67 1,274 3,176 35,332 122,660 1975 5,908 1,608 237 1,166 1,584 7,812 2,899 8,813 4,380 7,433 22,889 2,526 18,420 801 2,148 2,437 2,135 387 93,583 1,648 2,838 3,521 4,182 1,554 841 1,221 10,570 36,622 14,683 2,762 2,299 219 70 6,011 1,457 135 1,665 2,891 31,368 126,557 1974 5,390 1,423 204 1,753 2,084 7,306 3,084 9,521 4,253 8,459 25,384 3,007 20,221 1,075 2,628 2,376 1,721 425 100,314 1,191 2,348 4,544 3,666 2,138 1,364 775 10,063 33,455 10,233 2,120 1,837 271 168 6,052 628 37 1,751 1,976 30,420 115,037 1973 5,731 1,761 143 1,317 1,488 6,327 2,827 9,304 3,152 8,696 26,991 2,298 16,494 936 1,705 2,512 1,837 318 93,837 1,161 1,817 2,989 3,392 1,485 880 704 9,338 30,922 7,671 2,365 2,062 236 185 4,791 743 31 1,473 1,892 19,765 93,902 1972 5,006 1,138 195 1,265 1,872 5,503 2,013 9,187 3,388 6,925 24,284 2,337 17,662 729 2,136 1,731 1,733 688 87,792 1,585 2,693 4,205 3,227 1,374 638 429 16,039 28,312 8,621 2,486 1,414 163 214 4,524 716 41 1,416 2,016 24,667 104,780 1971 5,366 1,553 347 1,467 2,220 6,166 3,047 8,622 3,935 9,969 29,474 2,458 24,076 1,064 2,395 2,211 1,650 389 106,409 1,032 2,102 3,406 3,850 426 626 491 13,943 29,190 8,577 1,959 1,514 92 263 6,046 860 45 1,353 2,066 21,696 99,537 1970 7,286 1,403 180 2,961 1,558 5,798 3,252 10,148 2,973 7,728 25,789 2,748 22,030 794 2,480 1,859 1,611 482 101,080 1,449 1,117 2,483 2,784 452 1,893 354 17,636 29,894 9,360 2,195 2,053 172 37 5,268 611 46 624 1,753 27,885 108,066 1969 6,161 1,648 228 1,565 1,116 5,511 2,982 8,707 3,478 7,367 26,656 2,033 17,406 704 2,124 1,606 1,499 252 91,043 1,701 954 3,283 2,404 176 459 259 15,014 22,586 5,365 1,894 1,718 168 41 5,620 572 84 1,119 2,215 18,146 83,778 1968 4,450 1,163 157 1,576 1,429 5,522 2,551 7,884 2,613 5,815 21,977 2,138 12,069 483 2,449 1,578 1,187 314 75,355 1,571 912 1,347 1,444 170 463 293 9,733 17,711 3,756 848 762 33 194 3,561 376 32 571 875 12,182 56,834 1967 3,874 707 166 1,241 1,295 4,002 2,122 6,123 2,275 5,229 20,688 1,843 13,588 420 1,203 1,374 1,472 304 67,926 1,272 1,563 1,797 1,582 380 347 260 10,943 22,323 5,739 1,722 1,150 81 0 3,662 478 137 866 1,407 13,845 69,554 1966 3,388 829 197 1,331 820 4,115 1,462 6,787 2,128 4,525 16,712 2,169 7,688 492 1,165 1,608 843 197 56,450 1,424 639 1,454 1,387 121 245 351 11,112 20,480 2,932 1,337 934 56 110 2,958 192 45 676 1,578 11,730 59,761 1965 3,554 754 183 1,191 771 3,564 1,484 7,414 2,187 4,120 15,320 1,313 10,467 429 1,124 876 993 206 55,950 760 934 1,410 1,185 204 153 221 7,395 20,886 2,571 811 1,181 95 165 3,403 81 26 418 897 13,548 56,344 1964 3,129 706 91 758 812 2,980 1,407 5,548 1,594 3,462 12,699 1,735 7,317 382 1,398 1,017 1,086 93 46,214 828 1,394 1,328 1,480 570 364 224 10,327 23,631 4,536 1,422 1,255 207 88 5,357 265 33 298 1,799 18,257 73,663 Table 7. State and regional estimates of the number of duck stamp buyers that hunted American woodcock in the U.S., 1964-2001. Data from Martin (1979) and unpublished FWS administrative reports. Information was not available to reassign hunters from the District of Columbia to state of harvest. State or Region CT DE DC FL GA ME MD MA NH NJ NY NC PA RI SC VT VA WV Eastern Region AL AR IL IN IA KS KY LA MI MN MS MO NE ND OH OK SD TN TX WI Central Region 18 2001 1,353 245 53 352 1,045 3,168 1,382 1,510 1,083 1,046 3,493 1,209 5,277 187 1,292 920 719 212 24,546 400 135 1,655 1,353 772 97 891 2,130 12,961 6,923 917 1,304 396 160 3,221 325 95 903 1,720 7,322 43,680 2000 738 69 63 436 1,025 3,243 1,020 755 1,184 927 3,222 1,114 5,447 124 1,318 853 893 87 22,519 355 1,033 1,418 1,180 1,172 341 181 5,956 15,158 10,936 848 1,243 447 86 2,413 154 103 1,020 1,020 7,397 52,461 1999 1,305 0 50 346 586 3,791 855 2,033 956 1,282 4,193 1,529 5,885 221 1,477 941 1,564 119 27,131 323 1,637 1,417 1,304 721 930 839 5,617 17,539 8,197 177 998 196 111 2,503 520 0 1,145 734 8,670 53,578 1998 849 345 63 313 1,318 3,357 650 1,917 1,343 1,154 4,528 1,302 5,800 120 1,118 576 1,433 69 26,255 430 939 1,433 977 1,188 463 589 3,015 17,174 11,504 436 1,329 259 116 3,137 400 133 898 1,062 11,971 57,453 1997 1,470 286 74 598 758 2,903 607 1,772 1,031 1,269 4,439 1,192 5,753 174 1,045 950 1,305 142 25,767 374 542 1,846 1,010 1,244 275 311 2,816 16,789 11,455 639 1,408 400 75 3,600 250 76 1,016 1,453 11,389 56,968 1996 1,571 290 62 340 1,135 3,446 891 2,883 1,710 1,024 4,728 1,397 4,886 292 1,214 1,388 1,477 189 28,923 284 584 1,324 1,209 923 162 882 5,917 16,988 12,200 228 882 395 173 2,865 236 57 729 860 11,117 58,015 1995 1,929 401 48 189 837 3,009 1,161 2,992 1,511 1,298 4,837 1,017 5,670 269 950 1,300 1,004 158 28,581 331 241 1,831 1,207 1,060 273 909 5,654 18,503 11,745 574 774 185 225 2,617 378 85 899 318 11,115 58,924 1994 2,021 370 52 429 883 3,618 1,159 3,414 1,788 1,970 5,896 1,482 5,723 366 1,077 1,144 1,126 74 32,563 202 557 2,161 1,216 733 157 722 5,831 16,407 11,585 250 669 349 69 2,604 343 0 1,048 1,272 10,907 57,082 1993 1,916 226 60 463 1,217 3,718 1,125 3,617 1,652 2,378 5,122 1,172 6,464 262 1,153 840 944 71 32,397 250 113 2,118 1,424 489 160 569 7,988 16,547 7,933 330 1,051 282 269 3,080 405 0 536 969 11,060 55,573 1992 2,717 676 78 142 463 3,622 1,438 3,820 1,888 1,246 6,045 1,284 7,974 345 784 1,257 1,123 125 35,027 488 621 3,084 1,308 1,313 95 613 4,666 14,780 10,350 457 1,452 216 216 3,974 558 62 812 712 12,331 58,108 1991 3,016 620 45 549 838 4,016 1,494 4,195 1,666 1,793 6,244 1,855 7,701 420 951 1,485 1,427 104 38,464 337 914 3,575 1,125 1,292 140 704 5,981 16,445 11,735 417 962 177 201 3,075 333 129 717 1,288 12,787 62,334 1990 3,278 530 56 644 1,105 3,068 1,260 4,656 2,402 2,575 6,644 1,634 8,039 415 1,062 1,439 1,773 158 40,740 334 579 2,976 1,048 1,268 184 922 6,187 15,990 12,023 486 1,326 297 75 3,069 489 46 1,343 1,336 13,880 63,858 1989 2,958 392 33 551 1,036 3,043 1,365 4,816 2,029 2,672 7,901 1,752 7,550 404 1,457 1,299 1,540 197 40,995 730 573 2,913 1,499 1,506 338 910 8,478 14,296 14,212 676 1,796 260 262 4,189 620 125 837 1,519 14,353 70,092 1988 2,867 779 64 484 1,041 3,185 2,121 4,200 2,178 2,482 7,201 1,319 7,187 448 1,348 1,838 3,047 115 41,905 565 531 1,636 1,092 845 248 504 8,030 12,022 13,945 702 1,318 353 249 2,889 342 0 905 1,432 13,033 60,641 1987 3,687 731 74 1,075 1,311 4,195 2,842 5,117 2,433 2,743 9,716 1,809 10,055 567 1,912 1,977 2,317 203 52,764 605 1,638 3,078 1,722 1,957 812 746 14,792 14,563 14,076 828 2,252 507 41 3,350 502 137 1,276 2,264 17,084 82,230 1986 3,089 556 39 869 1,313 4,421 2,589 4,196 2,495 2,242 9,633 1,871 10,886 381 1,361 1,838 1,500 361 49,641 523 1,325 2,496 1,711 1,412 995 935 13,467 14,214 14,104 1,227 2,072 509 107 3,348 480 173 1,124 2,011 16,391 78,624 1985 3,390 438 49 658 1,312 4,196 3,147 4,733 2,380 3,274 9,418 1,885 9,546 478 1,266 1,629 2,168 213 50,182 731 1,404 2,533 1,349 1,622 600 698 13,907 13,888 10,524 1,116 2,141 374 177 3,879 668 27 1,010 1,502 14,874 73,024 1984 4,168 901 123 841 1,898 4,604 2,877 4,696 2,196 3,755 10,289 2,068 12,531 500 1,463 1,977 2,043 365 57,295 900 1,146 2,757 1,840 1,177 421 1,095 15,706 15,620 11,325 1,652 2,385 407 264 3,591 563 57 1,079 2,352 16,222 80,559 1983 4,385 691 0 1,057 1,901 4,459 2,512 4,129 2,460 4,411 10,515 2,066 12,816 619 2,054 1,828 1,011 543 57,459 675 1,094 2,938 1,529 1,546 415 858 20,526 12,642 10,414 1,394 1,565 737 175 4,188 647 132 765 1,931 15,052 79,223 Table 7. continued State or Region CT DE DC FL GA ME MD MA NH NJ NY NC PA RI SC VT VA WV Eastern Region AL AR IL IN IA KS KY LA MI MN MS MO NE ND OH OK SD TN TX WI Central Region 19 Table 8. Preliminary state and regional estimates of woodcock hunter numbers, days afield, and harvest from the 2001-02 and 2002-03 Harvest Information Program survey. Active woodcock hunters Days afield Harvest 2001-02 2002-03 2001-02 2002-03 2001-02 2002-03 Eastern region CT 1,800 ± 41% 1,600 ± 37% 7,500 ± 46% 9,200 ± 67% 3,600 ± 62% 4,500 ± 39% DE 400 ± 116% 500 ± 102% 5,200 ± 168% 900 ± 83% 200 ± 72% 500 ± 139% FL 2,400 ± 133% 1,000 ± 184% 14,600 ± 159% 2,000 ± 187% 9,500 ± 194% 100 ± 138% GA 3,600 ± 179% 2,600 ± 180% 40,500 ± 192% 5,600 ± 170% 11,000 ± 178% 600 ± 130% ME 11,900 ± 40% 5,000 ± 54% 64,900 ± 51% 18,500 ± 46% 48,100 ± 56% 18,600 ± 71% MD 700 ± 140% 600 ± 151% 1,500 ± 73% 1,100 ± 91% 1,700 ± 127% 600 ± 82% MA 1,200 ± 33% 1,100 ± 34% 5,700 ± 36% 6,000 ± 36% 2,500 ± 36% 3,500 ± 31% NH 2,000 ± 40% 1,500 ± 35% 9,900 ± 39% 7,500 ± 22% 6,700 ± 35% 5,600 ± 20% NJ 600 ± 66% 1,000 ± 69% 2,300 ± 23% 5,100 ± 86% 2,200 ± 30% 2,900 ± 57% NY 5,300 ± 37% 5,600 ± 36% 25,400 ± 41% 31,100 ± 47% 8,800 ± 55% 17,100 ± 62% NC 4,900 ± 154% 900 ± 67% 25,700 ± 147% 8,800 ± 104% 12,300 ± 126% 1,900 ± 132% PA 13,400 ± 45% 9,600 ± 44% 53,100 ± 52% 40,900 ± 57% 20,100 ± 52% 10,100 ± 40% RI 300 ± 88% 200 ± 82% 900 ± 101% 800 ± 73% 300 ± 63% 600 ± 83% SC 3,900 ± 92% 2,300 ± 129% 10,200 ± 107% 4,900 ± 122% 5,400 ± 171% 3,900 ± 163% VT 900 ± 39% 1,200 ± 45% 4,700 ± 36% 6,900 ± 55% 3,100 ± 28% 2,000 ± 31% VA 1,100 ± 128% 2,500 ± 86% 3,800 ± 108% 11,500 ± 96% 1,400 ± 30% 11,900 ± 176% WV 500 ± 82% 100 ± 21% 1,800 ± 100% 500 ± 30% 1,600 ± 73% 700 ± 42% Region na a na 277,800 ± 36% 161,200 ± 22% 138,100 ± 32% 84,800 ± 33% Central region AL 2,800 ± 109% 3,400 ± 93% 11,400 ± 146% 16,800 ± 95% 6,600 ± 191% 10,100 ± 109% AR 3,800 ± 131% 2,000 ± 172% 17,200 ± 166% 3,200 ± 113% 3,100 ± 132% 700 ± 112% IA 2,500 ± 78% 1,500 ± 103% 14,800 ± 113% 7,300 ± 134% 10,300 ± 128% 3,500 ± 130% IL 4,500 ± 81% 3,000 ± 90% 18,400 ± 82% 6,700 ± 86% 19,500 ± 112% 9,000 ± 111% IN 1,800 ± 106% 1,700 ± 113% 6,800 ± 118% 24,300 ± 172% 2,800 ± 96% 7,000 ± 160% KS 2,400 ± 110% 2,900 ± 96% 25,300 ± 113% 4,400 ± 111% 14,200 ± 138% 2,900 ± 137% KY 1,900 ± 174% 2,000 ± 126% 9,700 ± 171% 14,600 ± 150% 7,800 ± 171% 6,800 ± 141% LA 3,100 ± 139% 3,300 ± 148% 27,500 ± 155% 23,800 ± 166% 5,400 ± 59% 21,500 ± 138% MI 19,500 ± 23% 25,400 ± 18% 96,500 ± 24% 157,100 ± 37% 73,700 ± 33% 78,900 ± 26% MS 1,900 ± 133% 2,800 ± 187% 9,400 ± 154% 5,900 ± 179% 10,600 ± 129% 600 ± 64% MN 14,400 ± 49% 8,200 ± 66% 55,600 ± 47% 48,100 ± 94% 46,400 ± 71% 8,600 ± 31% MO 2,600 ± 101% 3,200 ± 125% 4,300 ± 89% 5,500 ± 115% 4,400 ± 114% 700 ± 40% NE < 50 ± 248% < 50 ± 155% 100 ± 73% 200 ± 72% 100 ± 99% 200 ± 74% OH 3,100 ± 135% 5,200 ± 108% 9,200 ± 93% 23,400 ± 137% 6,600 ± 87% 3,400 ± 43% OK < 50 ± 155% 2,500 ± 135% 200 ± 82% 6,300 ± 136% 100 ± 96% 2,600 ± 184% TN 200 ± 177% 4,200 ± 187% 700 ± 195% 4,800 ± 165% 700 ± 195% 8,000 ± 196% TX 10,400 ± 192% 28,700 ± 112% 12,800 ± 161% 67,000 ± 114% 5,300 ± 196% 38,300 ± 196% WI 14,800 ± 32% 17,100 ± 31% 68,700 ± 34% 57,500 ± 26% 33,700 ± 38% 32,000 ± 33% Region na na 388,600 ± 22% 476,800 ± 27% 251,400 ± 24% 235,100 ± 38% a Regional estimates of hunter numbers cannot be obtained due to the occurrence of individual hunters being registered in the Harvest Information Program in more than one state. 20 Appendix 1. History of framework dates, season lengths, and daily bag limits for hunting American woodcock in the U.S. portion of the Eastern and Central Regions, 1918-2002. Eastern Region Central Region Season Daily bag Season Daily bag Year (s) Outside dates length limit Year (s) Outside dates length limit 1918-26 Oct. 1 - Dec. 31 60 6 1918-26 Oct. 1 - Dec. 31 60 6 1927 Oct. 1 - Dec. 31 60 4 1927 Oct. 1 - Dec. 31 60 4 1928-39 Oct. 1 - Dec. 31 30 4 1928-39 Oct. 1 - Dec. 31 30 4 1940-47 Oct. 1 - Jan. 6 15 4 1940-47 Oct. 1 - Jan. 6 15 4 1948-52 Oct. 1 - Jan. 20 30 4 1948-52 Oct. 1 - Jan. 20 30 4 1953 Oct. 1 - Jan. 20 40 4 1953 Oct. 1 - Jan. 20 40 4 1954 Oct. 1 - Jan. 10 40 4 1954 Oct. 1 - Jan. 10 40 4 1955-57 Oct. 1 - Jan. 20 40 4 1955-57 Oct. 1 - Jan. 20 40 4 1958-60 Oct. 1 - Jan. 15 40 4 1958-60 Oct. 1 - Jan. 15 40 4 1961-62 Sep. 1 - Jan. 15 40 4 1961-62 Sep. 1 - Jan. 15 40 4 1963-64 Sep. 1 - Jan. 15 50 5 1963-64 Sep. 1 - Jan. 15 50 5 1965-66 Sep. 1 - Jan. 30 50 5 1965-66 Sep. 1 - Jan. 30 50 5 1967-69 Sep. 1 - Jan. 31 65 5 1967-69 Sep. 1 - Jan. 31 65 5 1970-71 Sep. 1 - Feb. 15 65 5 1970-71 Sep. 1 - Feb. 15 65 5 1972-81 Sep. 1 - Feb. 28 65 5 1972-90 Sep. 1 - Feb. 28 65 5 1982 Oct. 5 - Feb. 28 65 5 1991-96 Sep. 1 - Jan. 31 65 5 1983-84 Oct. 1 - Feb. 28 65 5 1997 *Sep. 20 - Jan. 31 45 3 1985-96 Oct. 1 - Jan. 31 45 3 1998 *Sep. 19 - Jan. 31 45 3 1997-01 Oct. 6 - Jan. 31 30 3 1999 *Sep. 25 - Jan. 31 45 3 2002 Oct. 1 - Jan. 31 30 3 2000 *Sep. 23 - Jan. 31 45 3 2001 *Sep. 22 - Jan. 31 45 3 2002 *Sep. 21 - Jan. 31 45 3 * Saturday nearest September 22. |
Original Filename | Am_woodcock_population03.pdf |
Date created | 2013-01-23 |
Date modified | 2013-03-06 |
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